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APR ^ 4 ?013

HARVARD

UNIVERSITY

VOL 14 (1)	2013 10 MARS
	SOMMAIRE
	Articles originaux - Original articles
P. Bail & A. Limpus	A new species of Ericiisa H. & A. Adams, 1858 (Gastropoda: Volutidae) from the bathyal Western Australian waters	1
W.G. Lyons &	New Caribbean Lamellilatirus (Gastropoda: Fasciolariidae:	5
M.A. Snyder	Peristerniinae) with a new record of a previously described species
G.T. Watters	Identity of Triton anomalus Hinds, 1844, and the description of a new eastern Pacific Bailya (Gastropoda: Buccinidae)	11
E. Rolân, R. Fernandez- A new species of Murchisonella (Heterobranchia,	17
Garcés & F. Rubio	Murchisonellidae) from Cuba
J. Pelorce	Deux nouvelles espèces de Columbellidae (Gastropoda: Neogastropoda) de l’île de Martinique, Antilles françaises	21
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P. Bail & A. Limpus

Novapex 14(1): 1-4, 10 mars 2013

A new species of Ericusa H. & A. Adams, 1858

(Gastropoda: Volutidae) MCZ

from the bathyal Western Australian waters LIBRARY

Patrice Bail

2, Square La Fontaine, 75016 Paris, France pat .bail® orange ir

Allan Limpus

6, MacKewen Street, Bundaberg, 4670 Queensland, Australia aslimpus@tpg.com.au

APR 2 4 2013

HARVARD

UNIVERSITY

KEYWORDS. Gastropoda, Volutidae, Southwest Australia, Ericusa naniforma sp. nov.

ABSTRACT. A new endemic species from the Southwest waters of the Western Autralian eontinental shelf is described here and compared with related species of the same genus Ericusa.

INTRODUCTION

During the early 1980s the Western Australian Fisheries Department designated an area off Rottnest Island where trawler operators eould test and try out their trawling equipment. This was done so as to préparé the boats before proceeding to their registered fishing grounds off Shark Bay and off Exmouth Gulf. As technology in the trawling industry progressed, this practice of tryouts eeased and further trawling operations came to an end.

During those trials many interesting species were brought up including numerous small Ericusa which differed from what was known, but were informally identified as ‘dwarf Ericusa papillosa ’.

The genus Ericusa is an Australian endemic genus, inhabiting the Southern waters below the 30° parallel, characterized by a solid shell, fusiform, medium to large, with a non-coronated elongate spire and regularly convex whorls. The protoeonch is globose, deviated at 45° from the axis of the shell and frequently with the initial portion protruded. It includes five species and one subspecies: Ericusa fulgetra (Sowerby I, 1825), Ericusa papillosa papillosa (Swainson, 1822), Ericusa papillosa kenyoniana (Brazier, 1898), Ericusa sericata Thornley, 1951 , Ericusa sowerhyi (Kiener, 1839), and the new species we describe here which has been overlooked for many years.

Abbreviations

WAM: Western Australian Muséum, Perth. ca: circa.

SYSTEMATICS

Family VOLUTIDAE Rafinesque, 1815 Subfamily ZIDONINAE A. & H. Adams, 1853 Tribe Livoniini Bail & Poppe, 2001 Genus Ericusa A. & Fl. Adams, 1853

Type species: Voluta fulgetrum Sowerby 1, 1825

Ericusa naniforina sp. nov.

Figs 1-17

Ericusa papillosa - Wilson, 1994: p. 123, pi. 23, figs 3A-B (only) [(not Ericusa papillosa (Swainson, 1822)].

Type material. Holotype in WAM, n° Si 1656. Length: 62.2 mm, width: 34.2 mm.

Paratypes. Paratype I from off Bunbury, no depth given, 65.6 mm, A. Limpus collection; Paratype II from SW Rottnest -300 m, 65.8 mm, P. Bail colleetion; Paratype III from off Rottnest, -280 m, 60.1 mm, J. Phillip eollection.

Other material. Four specimens in A. Limpus eollection: 64.9 mm, 62.2 mm, 60.9 mm, 59.5 mm, trawled -400/450 m off Southwest Rottnest island; two in P. Bail collection: 63.6 mm, 60.2 mm, trawled -300 m off SW Rottnest; one in J. Philipp collection: 60.1 mm, trawled - 280 m off Rottnest; two in P. Ignoty collection: 65.5 mm, 68.0 mm, trawled -350 m off Cape Leeuwin; one in E. Monnier collection: 61. 4mm trawled -350 m off Bunbury; in WAM collection: 1. n° S80349, ca 64.0 mm, [almost complété specimen] NW of Bunbury, 33°S - 1 14.37°E. HMAS

"Diamantina", 17-3-1972, depth 219-221m, 2.

S. 80347 [2 fragments of body whorl] NW of Bunbury 33°S - 114.38°E, HMAS "Diamantina", 15/3/1972, depth 256m, 3. SI 1666 [3 fragments] off Three Rocks -31°6036 S - 114°9850 E - to 3r6103 S - 114.9790°E., RV "Southern Surveyor" depth 275/327m, 19-11-2005, 4. S80348 [fragment only] off Lancelin, 3L04°S - 113.5°E, HMAS

"Diamantina", depth 256m, 23.3.1972, 5. S80350 [two fragments] NW of Rottnest Is. 3L45°S - 115.02°E, HMAS "Diamantina", depth 265-276m, 18-3-1972.

P.BA1L& A.LIMPUS

A nevv species of Ericusa from Western Australia

Type locality. Off Abrolhos Islands, 28° 48' S - 113° 41' S to 28° 50' E - 1 13° 42' E. Depth: -416 to -431 m. CSIRO RV "Southern Surveyor". Dec. 2005.

Range. From Houtman Abrolhos [vvest of Geraldton] to Cape Leeuwin, a rather limited range of 700 km.

Habitat. Most specimens hâve been travvled between 350-450 m deep. Substrat not given, but most specimens are stained by a rusty layer.

Description. Adult shell very small for the genus, rarely exceeding 65 mm, solid, ovate fusiform vvith a tapered spire. Surface semi-glossy. Protoconch bulbous, protruded, of 2.5 smooth whorls deviated at 45° from axis of shell, with an average diameter of 7 mm. Transition protoconch - teleoconch gradually marked by occurrence of faint axial striae. Teleoconch of 2.75 gently convex whorls, sculptured by straight; keel-like axial ridges, extending onto the two last whorls, mostly prominent and counted ca 45-55 on the last whorl, very attenuated and more numerous on some specimens. Spiral sculpture of close-set faint minute threads extending onto the whole teleoconch, slightly attenuated on the anterior half. Suture simple. Aperture semi-lunar forming an average of 0.72 of total shell length. Outer lip convex, the posterior half thickened and reflected, forming defined canal at adapical end of aperture. Columella almost straight, bearing 4-5 oblique plaits, the adapical the smaller. Fasciole absent. Siphonal notch shallow and wide. Background color yellowish-cream overlain by light pattern of irregular orange-beige blotches forming two wide spiral bands on mid body whorl and on the anterior third, with remnants of marks on the subsutural zone. Protoconch, aperture and columellar plaits beige. Frequent incrustations of rusty deposits on the crest of axial ribs and on the spiral threads

giving irregular revolving rusty bands on some specimens.

Animal. Unknown.

Discussion. We disregard the other members of the genus Ericusa jïdgetra, Ericusa sericata and Ericusa sowerbyi which are of large size, different shape and well-differentiated pattern which preclude any confusion.

By its very spécifie characters such as the very small adult size and ridged surface, E. naniforma must be only compared with the ridged specimens of its closely related species E. papillosa.

Two populations of sculptured E. papillosa occur along the species range:

The first, E. papillosa kenyoniana represents the eastern population extending from Coffs Harbour to the east coast of Tasmania and the coast of Victoria east of Fakes Entrance. Generally considered a valid subspecies, it differs from the Southern type species E. papillosa papillosa by an elongate shape and the presence of solid axial ribs on the whole teleoconch. No confusion with E. naniforma is possible because of its large size, above 120 mm, a depressed rounded protoconch, a shiny surface with a conspicuous tent- like pattern.

The second ridged form of E. papillosa occurs here and there ail along the species range from Bass Strait to Augusta, especially at a deeper level (350-450 m) than the type species (50-150 m). It is characterized by a smaller size, a squat shape and by irregular axial ridging on the teleoconch.

There is an obvions resemblance between this form and E. naniforma, but différences are constant (see table) and the shells are unmistakable (plate 2, figs 18- 21):

	Ericusa naniforma	ridged dwarf Ericusa papillosa
protoconch	bulbous, protruded	rounded, partly immersed
spire	tapered	large, triangular
shape	elongate ovate	rhomboid
outer lip	convex	slightly flattened in the middle
sculpture	fine, regular, keel-like	coarse, irregular, rounded
pattern	hardly visible	well-defined
siphonal notch	relatively nairow	relatively broad

Figures 1-17 Ericusa naniforma sp, nov.

1-3. Off Abrolhos Islands, 28° 48' S -1 13° 41’ S to 28° 50' E - 1 13° 42' E, 4 16-431 m, holotype WAM SI 1656, 66.1 mm.

4-5. Off Rottnest, 64.9 mm A.L. collection; 6-7. Off Rottnest, 60.9 mm A.L. collection; 8-9. Off Rottnest, 60.1 mm J. Ph.. collection, paratype III; 10-11. Off Rottnest, 65.8 mm P. B. collection, paratype II; 12-13. Off Rottnest, 63.6 mm P. B. collection; 14-15. Off Bunbury, 65.6 mm A.L. collection, paratype I; 16-17, Off Rottnest, 62.2 mm A.L. collection.

2

P. Bail & A. Limpus

Novapex 14(1); 1-4, 10 mars 2013

3

P. Bail & A. Limpus

A new species of Ericusa from Western Australie!

Reiiiarks. This speeies E. naniforma présents very tew variations, differing only in the number or strength of axial ridges. In most specimens, the light pattern is hardly visible, covered by a thin rusty layer. The presence of ridges in E. papillosa is connected vvith the depth ail along its range and seems linked with a reduced size along the South Australian coast. These shell characters show évident similarities with E. naniforma which probably is subject to the saine ecological constraints. These two species are very likely related.

Nevertheless, a biogeographical barrier around the Cape Leeuwin séparâtes them and no intermediate shells hâve been found there up to now, which allows one to suppose the genetic flow to be interrupted, leading to a séparation at spécifie rank with constant and unmistakable différences.

The currents which flow strongly around Cape Leeuwin make a barrier by themselves and this would

be why very few west coast species extend further to the east except several shallower water species of which very few are known. To understand this phenomenon, one can stand at Cape Leeuwin and see the flow of the currents as they swirl around the cape, where it is the meeting of the Circum Polar Antarctic current which flows in a constant easterly direction, and the Leeuwin Current which flows down the W.A. coast. The Leeuwin Current then is forced eastwards, before dissipating in the Great Australian Bight , while the Arctic current continues eastwards to Bass Strait and Tasmania. However, the possibility of sympatry between naniforma and dwarf papillosa cannot be excluded since specimens possibly similar to E. naniforma hâve been reported found in the far western Great Australian Bight (H. Morrison, pers. com.).

Etymology. Related to its very small size.

Figures 18-21

18 & 20. Ericusa naniforma Bunbury, P. B. collection, A.L. collection, 56.5 mm.

Acknowledgments. Our appréciation goes to Arron Cosgrove-Wilke of the Western Australian Muséum for his assistance and the photographs of the WAM specimens, also to Hugh Morrison for his work on our behalf and for relevant information. We also thank John Phillips and lan Mattiske for allowing us to photograph specimens from their collections, and

65.6 mm; 19 & 21. Ericusa papillosa South Australia,

Peter Ignoti for supplying photographs of specimens from his collection.

REFERENCE

Wilson, B. 1994. Australian Marine Shells. Vol. 2. Odyssey Publishing, Kallaroo: 1-370.

4

W.G. Lyons & M. A. Snyder

Novafex 14(1): 5-10, 10 mars 2013

New Caribbean Laimllilatirus (Gastropoda: Fasciolariidae: Peristerniinae) with a new record of a previously described species

William G. LYONS

4227 Porpoise Drive SE, St. Petersburg, Florida 33705 USA \v.lyons9@knology.nel Corresponding author

Martin Avery SNYDER

Academy of Natural Sciences of Drexel University, 1900 Benjamin Franklin Parkway, Philadelphia, Pennsylvania 19103 USA and Muséum national d'Histoire naturelle, Paris

dr.martin.snyder@gmail.com

KEY WORDS. Mollusca, Gastropoda, Fasciolariidae, Peristerniinae, LameUilatinis, Recent, Caribbean Sea, new species

ABSTRACT. Four new species of Lamellilatirus Lyons & Snyder, 2008, L. eburneiis, L. lamyi, L. dominiqiiei and L. siinderlandoriim, are described and distinguished from each other and from the type species, L. ceninudus (Dali, 1889). The new species are from varions locations in Guadeloupe, Martinique, Venezuela, Colombia and Honduras in the Southern Caribbean Sea. The range of L. ceramidus, previously known from Barbados, Colombia, Panama and Nicaragua, is extended to Puerto Rico.

INTRODUCTION

LamelUlatirus Lyons & Snyder, 2008, was introduced as a monotypic genus to contain Fusus ceramidus Dali, 1889, whose radula was revealed by Bullock (1968) to be peristerniine, not fusinine. Unusual features of the shell and radula dictated that a new genus be erected to contain the species. Recent studies of specimens in several private collections hâve revealed four additional undescribed species which are appropriately placed in LamelUlatirus .

Material and Methods

Ail specimens were acquired from commercial shell dealers or amateur collectors. Animais had been removed and discarded but opercula still accompanied many shells. Some shells had passed through several hands before we obtained them, but most species were represented by several lots obtained from different sources, and data with the lots are in sufficient agreement to instill confidence in general accuracy. Measurements are reported in metric units. Depths of collection, sometimes initially recorded in feet (ft) or fathoms (fm), are converted to meters (m). Shell sizes are reported in millimeters (mm), measured to nearest 0.1 mm with electronic digital calipers; single measures are of shell length (si; = height), whereas two measures signify shell length and width (si x w) or a range of shell heights (si - si); Iv dénotés live- taken specimens, dd empty shells.

Abbreviations

Specimens were examined in private collections of Kevan and Linda Sunderland (SC) of Plantation, Florida and the senior author (LC). Dominique Lamy of Guadeloupe, French West Indies, provided several important specimens which he donated to institutional collections. Abbreviations for institutional catalogue numbers are: ANSP, Academy of Natural Sciences of Drexel University, Philadelphia, Pennsylvania; BMSM, Bailey-Matthews Shell Muséum, Sanibel, Florida; MNHN, Muséum national d'Histoire naturelle, Paris; UF, Florida Muséum of Natural History, Gainesville; and USNM, United States National Muséum of Natural History, Washington, D.C.

SYSTEMATICS

Family FASCIOLARIIDAE Gray, 1847 Subfamily PERISTERNIINAE Tryon, 1881 Genus LamelUlatirus Lyons & Snyder, 2008 Type species by original désignation: Fusus ceramidus Dali, 1889, Recent, Barbados.

Remarks. LamelUlatirus ceramidus (Figs 1 , 2) was previously known by specimens from the Southern Caribbean Sea (Barbados, Colombia, Panama and Nicaragua) in depths of 73-220 m (Lyons & Snyder, 2008). The species has a polished white shell of moderate size with a somewhat swollen spire, broad

5

W.G. Lyons & A. Snyder

New Caribbean lumieUilatirus

axial ribs, reduced or obsolète spiral cords, reduced columellar plicae, internai lirae entire or interrupted as beaded pustules, a slender siphonal process, and numeroLis axial lamellae on the suturai ramp. The radula was figured by Lyons & Snyder (2008: fig 3a), after Bullock (1968: pl. 8, fig 7). Although Latiriis- like, the laterals hâve distinctive rounded cusps near the inner margin, whereas most peristerniines hâve sharply pointed cusps. The generic diagnosis is modified here to accommodate new species with relatively more slender spires, more prominent spiral cords, less prominent suturai lamellae and more colorful shells.

The only other Caribbean généra of Peristerniinae with conspicuous suturai lamellae are Polygona Schumacher, 1817 [type species Polygona jïisiformis Schumacher, 1817, = Murex infundibulum Gmelin, 1791, Recent, Caribbean Sea, by monotypy, fide Vermeij & Snyder (2006: 419, 420)] and some Piistulatirus Vermeij & Snyder, 2006 (type species Latirus mediamericanus Hertlein & Strong, 1951, Recent, tropical eastern Pacific, by original désignation). However, both Polygona and Pustulatirus species generally live in shallower water and their columellar plicae remain conspicuous in mature shells. Shells of Polygona are heavier, more compact, and usually hâve much shorter siphonal processes. Mature shells of Pustulatirus hâve a serrate outer lip formed by extensions of interspaces between spiral cords of the body whorl, whereas shells of Lamellilatirus lack labral serrata.

We encountered one specimen that extends the range of L. ceramidus northward in the eastern Caribbean. The new record is a 59.0-mm shell taken dead in a fish trap off Mona Island, Puerto Rico, depth 220 fm [403 m] (LC; Figs 3, 4). The specimen represents new depth and size records for L. ceramidus. However, the Shell was probably carried into the trap by a crab and the specimen may hâve lived in a lesser depth.

Lainellilatinis eburneus n. sp.

Figs 5-7, 24

Type material. Holotype (43.4 x 16.2 mm), dd, off Isla San Bernardo, Colombia, trawled, depth 150 m, ANSP 450383.

Other material. 1 (24.9 mm), dd, off Islas de Los Testigos, Venezuela, dredged, 195 m, LC.

Distribution. Venezuela and Colombia, Southern Caribbean Sea; depth range 150-195 m.

Description. Shell of moderate size for genus, to 43.4 X 16.2 mm, fusiform, slender, with about 10 Vi whorls, elevated spire, elongate siphonal process and scattered suturai lamellae. Protoconch of about 2 14 whorls, first whorl globose, tip immersed, second whorl not much wider than first, with slightly convex sides and about 4 faint axial riblets near terminus. Teleoconch of about 8

convex whorls separated by deeply impressed suture and sculpted with prominent axial ribs and spiral cords and with sparse axial lamellae scattered on rudimentary suturai ramp; axial ribs of ail whorls broad, node-like, extending from anterior suture to posterior suturai ramp; 5 spiral cords on early whorls, of nearly equal size on whorl 1; middle 3 cords becoming stronger on whorls 2-4; ail cords strong on whorls 5-7. Body whorl somewhat shouldered, with 2 or 3 very prominent cords at periphery; 2 or 3 more cords diminishing in strength apically and 3 or 4 cords diminishing in strength toward base; surface of spire and body whorl with many fine, axially aligned microlamellae and as many as 6 spiral threads in interspaces between cords; about 12 oblique cords, some separated by fine threads on base and siphonal process. Aperture subovate; outer lip highly arched, thin, crenulated at edge by spiral cords and interspaces of body whorl, inner side of lip with 8 or 9 smooth, emergent lira extending to lip edge; columella gently curved, with about 2 faint folds near anterior end; pariétal shield smooth; siphonal canal slender, smooth within, twisted near middle to form small pseudoumbilicus near anterior end. Protoconch and exterior and interior of fresh shell pure white; older Shell fading to cream. Operculum and radula unknown.

Etymology. From the Latin eburneus, an adjective meaning “ivory-colored,” referring to the pure white shells whose colors fade to cream like old ivory as they âge.

Remarks. Shells of L. eburneus differ from those of L. ceramidus by having a relatively taller, more slender spire and more scattered and sparse axial lamellae. The shell surface of L. ceramidus is nearly smooth, whereas that of L. eburneus has strong spiral cords on ail whorls; the two or three large cords Crossing the tops of the axial ribs characterize the species. Différences with other species are discussed in accoLints that follow.

The 24.9-mm Venezuelan shell is too old and wom to be included as a type.

Laine llila tir us lamyi n. sp.

Figs 8-12,25

Type material. Holotype (38.3 x 15.2 mm), off Pointe de la Caravelle, east coast of Martinique, French West Indies, depth 250 m, MNHN 25714; paratype (20.6 mm), same data as holotype, ANSP 450384.

Distribution. Known only from type locality off Martinique, French West Indies, depth 250 m.

Description. Shell of moderate size for genus, to 38.3 X 15.2 mm, fusiform, slender, with about 9 whorls, prominent axial ribs and spiral cords, and abundant axial lamellae. Protoconch of 2 elevatecj whorls, first

6

W. G. Lyons & M. a. Snyder

Novapex 14(1): 5-10, 10 mars 2013

vvhorl smooth, rounded, tip immersed; second whorl not much vvider than first; sides slightly convex, vvith 3-4 very faint axial riblets near terminus. Teleoconch of 7 vvhorls, each with 6 or 7 rounded, node-like, seldom aligned axial ribs; whorls 1-4 each vvith 4 spiral cords; 2 middle cords Crossing ribs stronger than others; cord bordering anterior suture increasing in strength tovvard anterior vvhorls; vvhorls 5 and 6 with 5 cords, 3 anterior cords stronger. Body whorl with about 1 1 cords of varions strengths; 3 cords at and anterior to periphery more prominent than others; surface of spire and body whorl with many axially aligned micro-lamellae and (sometimes) with 1-3 fine spiral threads between cords. Suture deeply impressed, undulant in accord vvith ribs and interspaces; suturai ramp bearing many prominent scale-like lamellae on middle vvhorls (3-5); lamellae less prominent on anterior vvhorls; base and siphonal process slender, elongate, vvith 8-10 oblique cords of varions strengths extending to truncate tip. Aperture subovate; outer lip thin, highly arched, slightly crenulate, inner side bearing 9-11 low, straight, sometimes discontinuons lirae; several anteriormost lirae beaded. Columella nearly straight anteriorly, arched posteriorly, with small node-like calions on pariétal shield and faint ondulations suggestive of folds near calions at anterior end. Siphonal canal slender, straight, smooth within, with lamellate inner edge forming distinct pseudoumbilicus near tip. Shell exterior light orange- brown, with lighter hue extending in vague band across ribs of spire; protoconch and shell interior white. Operculum and radula unknown.

Etymology. The name honors M. Dominque Lamy, Baie-Mahault, Guadeloupe, whose collections hâve enriched our knowledge of Caribbean Mollusca. M. Lamy provided specimens of two species treated in this paper, including both specimens of the species here described.

Remarks. Brown shells of L. lamyi distinguish that species from L. ceramidus and L. eburneus, which hâve white shells. Spiral cords that cross the ribs of L. lamyi are stronger than those of L. ceramidus but vveaker than those of L. eburneus. Suturai lamellae are less densely packed on L. lamyi and L. eburneus than on L. ceramidus but are arrayed over more of the suturai ramp on L. lamyi than on L. eburneus. The larger shell (holotype) of L. lamyi has beaded lirae within the aperture, a feature shared with L. ceramidus but apparently not with some other congeners.

The outer lip of the holotype is thin and not fully developed, suggesting that the specimen may be immature and the species may attain somewhat greater size.

Lamellilatirus dotniniquei n. sp.

Figs 13-17,26

Type material. Holotype (29.3 x 1 1 .1 mm), off Cabo

de la Vêla, Colombia, trawl, depth 200 m, ANSP 450385; paratype (21.4 mm). Saint François, eastern Guadeloupe, French West Indies, depth 200 m; MNHN IM-20 12-25.

Distribution. Guadeloupe and Colombia, Southern Caribbean Sea, in 200-m depths.

Description. Shell small for genus, to 29.3 x 11.1 mm, fusiform, slender, vvith about 9 whorls, elevated spire and elongate siphonal process. Protoconch of 2 elevated whorls; first whorl smooth, rounded, with immersed tip; second whorl no wider than first, with 4-5 faint axial riblets near terminus. Teleoconch of 7 convex whorls separated by deep suture and sculpted with prominent axial ribs and spiral cords, and with fine axial lamellae on suturai ramp; first whorl with about 8 small axial ribs, thereafter with 6 prominent, broad, elevated ribs on each whorl, ribs aligned from whorl to whorl on spire, not aligned on body whorl; 3 equal-sized spiral cords on whorl 1 , increasing to 4 on whorls 2-4 and to 5 on whorls 5 and 6, 3 middle cords stronger than others; 8 cords on body whorl, 4 Crossing tops of axial ribs most prominent; many very fine spiral threads and axially aligned micro-lamellae between cords of spire and body whorl. Suture impressed, undulant in accord with ribs and interspaces; suturai ramp bearing many very thin lamellae overlying faint spiral threads; base and siphonal process slender, elongate, with 8-12 narrow, oblique cords extending to rounded tip. Aperture subovate; outer lip crenulated by cords and interspaces of body whorl; inner side with a few thin, straight lirae. Columella arcuate, smooth except for small node on pariétal shield and 2 faint, oblique folds near callous at anterior end; siphonal canal slender, straight, smooth within. Protoconch and shell exterior light brown or tan; larger cords of body vvhorl darker reddish brown; interior white. Operculum elliptical, gray-brown, with anterior nucléus and many fine, concentric grovvth incréments on outer surface. Radula unknown.

Etymology. The name honors M. Dominque Lamy, Baie-Mahault, Guadeloupe, who provided one of the two specimens used in the description of the species.

Remarks. Shells of L. dominiquei, like those of L. lamyi, are basically brown but lack the lighter hued peripheral band of the latter species; instead, the three or four strongest cords Crossing the axial ribs of L. dominiquei are dark reddish brown, especially on the body whorl, providing a conspicuous contrast vvith the rest of the shell. Compared to shells of L. lamyi, those of L. dominiquei are also smaller, with a relatively more slender spire and much finer, less numerous suturai lamellae. Both specimens of L. dominiquei hâve thin, immature lips, and it seems likely that fully adult specimens may be larger. However, the number of teleoconch whorls (seven) of L. dominiquei is

7

W.G. Lyons & A. Snyder

New Caribbean LamellUatirus

similar to that of the larger immature shell of L. lamyi, so L. dominiqiiei may ultimately prove to be a smaller species.

The brown shell of L. dominiqiiei séparâtes that species from L. ceramidiis and L. eburneiis. In addition, L. ceramidiis bas more prominent and abundant suturai lamellae, and its spire is more swollen, the ribs are broader, and the cords more subdued.

LamellUatirus sunderlandorum n. sp.

Figs 18-22,27

Type material. Holotype (33.3 mm), dd, off Pulpit Rock, Mangrove Bight, Roatân, Honduras, trap, 270- 280 m, ANSP 450386. Paratypes: 1 (32.2 mm), dd, same data as holotype, MNHN IM-20 12-26; 2 (35.3 and 18.0 mm), dd, same data as holotype, SC; 2 (27.6 mm, Iv; 27.2 mm, dd), off Flores Bay Church, Roatân, trap, 124 m, SC; 4 (19.2 and 11.2 mm, Iv; 16.6 and 15.3 mm, dd), off Roatân, shell traps, 200-300 ft [61- 91 m], SC; 1 (23.7 mm), Iv, off Roatân, baited shell trap, 300 ft [91 m], UF 456795; 1 (18.7 mm), Iv, same data, BMSM 17934; 3 (20.8, 17.6 and 15.5 mm), dd, same data, LC; 1 (35.0 mm), dd, off north coast of Roatân, dredge, 350 m, SC.

Distribution. Known only from rather deep water (61-350 m) off the northern coast of Honduras, western Caribbean Sea.

Description. Shell of moderate size for genus (to 35.3 X 12.4 mm), fusiform, slender, of about 9 whorls, with tall spire and slender siphonal process. Protoconch of about 2 elevated whorls; first whorl smooth, rounded, with immersed tip; second whorl slightly wider, with nearly straight sides and about 4 very faint axial riblets near terminus. Teleoconch of about 7 whorls bearing prominent axial ribs and low spiral cords; suture narrow, impressed, undulant in accord with ribs and interspaces. Axial ribs 7, high, broad, node-like, bordered anteriorly by suture and posteriorly by short suturai ramp; 3 spiral cords of similar size on whorls 1-3; cords thereafter altemating larger and smaller in size and gradually increasing in number to 10 on penultimate whorl and about 15 on body whorl; strongest cord at periphery atop ribs. Suturai ramp

marked with several spiral threads Crossing reduced suturai lamellae, creating reticulate pattern. Base and siphonal process elongate, slender, bearing about 1 1 low, oblique cords and terminating at rounded tip. Aperture subovate, constricted posteriorly by well- developed pariétal node and anteriorly by prominent node opposite node at junction of columella and siphonal canal; outer lip arched, generally smooth at edge, inner side bearing 9-14 straight lirae not extending to lip edge; inner lip thin, elevated on largest specimens, extending from pariétal région to canal. Columella nearly straight anteriorly, arched posteriorly, bearing 3-4 oblique plicae which diminish to obscure folds on largest shells; siphonal canal straight, smooth within; larger shells (> 32 mm si) bordered on inner side by lamellate extension of inner lip which extends to tip, creating chink-like pseudoumbilicus on siphonal process. Exterior of shell orange to orange-brown; interior white. Operculum brown, ovate, vaguely drop-shaped, tapering to anterior nucléus; outer surface bearing many fine concentric growth incréments. Radula unknown.

Etymology. The species is named for Kevan and Linda Sunderland of Plantation, Florida, who provided many of the specimens we examined and whose collection has proved invaluable in this and other studies.

Remarks. Suturai lamellae of L. sunderlandorum shells are much reduced as compared to those of their congeners, but close examination reveals fine, low but distinct lamellae on the suturai ramp. Other features consistent with LamellUatirus are the morphology of the protoconch (Figs 23-27), the tall spire, the generally slender shell, the relatively long, narrow siphonal process, the non-semate edge of the outer lip, and the progression from distinct columellar folds in young shells to obscure folds on the largest shells. The prominent node at the anterior end of the aperture and an opposing node at the junction of the columella and siphonal canal, présent only on the largest L. sunderlandorum (35.0 and 35.3 mm si), are features that also occur on larger shells of L. ceramidus.

Figures 1-22. Scale bar: 5.0 mm.

1-4. LamellUatirus ceramidus (Dali, 1889). 1-2. Lectotype, 46.2 mm, Barbados, depth 134 m, USNM 87069; 3- 4. Off Mona Island, Puerto Rico, 403 m, LC, 59.0 mm.

5-7. LamellUatirus eburneiis n. sp. 5-7. Holotype, 43.4 mm, ANSP 450383, off Isla San Bernardo, Colombia, 150 m.

8-12. Uimellilatirus lamyi n. sp. 8-10. Holotype, 38.3 mm, MNHN 25714, off Pointe de la Caravelle, Martinique, 250 m; 11-12. Paratype, 20.6 mm, ANSP 450384, same data as holotype.

13-17. LamellUatirus dominiquei n. sp. 13-15. Holotype, 29.3 mm, ANSP 450385, off Cabo de la Vêla, Colombia, 200 m; 16-17. Paratype, 21 .4 mm, MNHN IM-2012-25, Saint François, Guadeloupe, 200 m.

18-22. LamellUatirus sunderlandorum n. sp. 18-20. Holotype, 33.3 mm, ANSP 450386, off Pulpit Rock, Mangrove Bight, Roatân, Honduras, 280-270 m; 21-22. Paratype, 32.2 mm, MNHN IM-2012-26, same data as

holotype.

8

W . G . LYONS & M . a . SN YDER

NovAf^EX 14(1); 5-10, 1 0 mars 20 1 3

9

W.G. Lyons & A. Snyder

New Caribbean Lamellilatinis

Shells of L. simderlandorum are orange, whereas those of L. ceramidus and L. eburneiis are white and those of L. dominiqiiei and L. lamyi are brown, the latter with a lighter-hued band at the periphery. Spiral cords of L. simderlandorum are much smaller than those of L. ebiirneus, L. lamyi and L. dominiquei . Like L. ceramidus, spiral cords alternate large and small, but ail cords of L. sunderlandoriim are proportionally smaller than those of L. ceramidus.

Lamellilatinis sunderlandoriim also resembles a new species of Pustulatirus with which it sometimes

occurs off the Honduran coast (Lyons and Snyder, in prep.) Both shells are orange, but L. sunderlandorum is longer and more slender than the new Pustulatirus, which has less rounded whorls, a shallower suture lacking lamellae, and a serrate edge on at least the anterior portion of its outer lip.

Both the inner and outer lip of the largest paratype (35.3 mm si) of L. simderlandorum are chipped and the body whorl bears a conspicuous hole similar to some we hâve seem on shells ex pisce. The 35.0-mm paratype also has a severely chipped outer lip.

Figures 23-27. Protoconchs; scale bar: 1.0 mm.

23. Lamellilatirus ceramidus (ANSP 450455); 24. Lamellilatinis eburneus, holotype (ANSP 450383); 25. Lamellilatinis lamyi, holotype (MNHN 25714); 26. Lamellilatirus dominiquei, holotype (ANSP 450385); 27. Lamellilatirus sunderlandorum, holotype (ANSP 450386).

Acknowledgements

We thank Dominique Lamy and Kevan and Linda Sunderland for making available many of the specimens examined in this study. Amanda Lawless, ANSP, photographed the specimens and prepared the plates. Helpful review comments were provided by Drs. E. F. Garcia and M. G. Harasewych.

REFERENCES

Bullock, R. C. 1968. The fasciolariid généra Latirus, Dolicholatirus, and Teralatirus (Mollusca: Gastropoda) in the western Atlantic. M. Sci. Thesis, University of Maine, Orono, August 1968. 109 pp.

Dali, W. H. 1889. Reports on the results of dredging, under supervision of Alexander Agassiz, in the Gulf of Mexico ( 1 877-78) and the Caribbean Sea ( 1 879-80), by the U. S. Coast Survey steamer "Blake", Lieut.-Commander C. D. Sigsbee, U. S. N., and Commander J. R. Bartlett, U. S. N., commanding. XXIX. Report on the Mollusca. Part 11. Gastropoda and Scaphopoda. Bulletin of the Muséum of Comparative Zoology at Harvard College 18: 1-492, pis. 1-40.

Gmelin, J. F. 1791 . Vermes. In: Caroli a Linnaei, systema naturae per régna tria naturae. Editio décimal tertia, Aucta, Reformata, Lipsiae. 1(6): 3021-3910.

Gray, J. E. 1847. A list of the généra of Recent

Mollusca, their synonyma and types. Proceedings of the Zoological Society of London 15(2): 129- 219.

Hertlein, L. G. & Strong, A. M. 1951 . Eastern Pacific expéditions of the New York Zoological Society. XLIIl. Mollusks from the west coast of Mexico and Central America. Part 10. Zoologica 36: 67- 120, pis. 1-11.

Lyons, W. G. & Snyder, M. A. 2008. New généra and species of Peristemiinae (Gastropoda: Fasciolariidae) from the Caribbean région, with comments on the fasciolariid fauna of Bermuda. The Veliger 50: 225-240.

Lyons, W. G. & Snyder, M. A. In prep. The genus Pustulatirus Vermeij and Snyder, 2006 (Gastropoda: Fasciolariidae: Peristemiinae) in the western Atlantic, with descriptions of three new species. [Submitted]

Schumacher, C. F. 1817. Essai d’un nouveau système des habitations des vers testacés, avec XXll planches. Schultz, Copenhagen. 287 pp., 22 pis.

Tryon, G. W. 1881. Manual of conchology , structural and systematic, with illustrations of the species. Volume 3. Tritonidae, Fusidae, Buccinidae. Philadelphia. 310 pp., 87 pis. [5-64, 1880; 65-310, 1881].

Vermeij, G. J. & Snyder, M. A. 2006. Shell characters and taxonomy of Latirus and related fasciolariid groups. Journal of Molluscan Studies 72: 413-424.

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G.T.Watters

Novapex 14(1); 11-15, 10 mars 20 13

Identity of Triton anotnalus Hinds, 1844, and the description of a new eastern Pacific Bailya (Gastropoda: Buccinidae)

G. Thomas WATTERS

Department of Evolution, Eeology and Organismal Biology Ohio State University, Columbus, Ohio, 4312, USA Watters.l @osu.edu

KEYWORDS. Eastern Pacific Océan, Central America, Gastropoda, Buccinidae, Bailya cidaris n. sp.

ABSTRACT. A lectotype is designated for Triton anomalus Hinds, 1844, now Bailya anomala. That species is redescribed and Bailya cidaris n. sp. is described from western Panama.

INTRODUCTION

Examination of specimens of the buccinid genus Bailya from western Panama bas revealed the presence of two species currently placed under Bailya anomala (Hinds, 1844). Two names hâve been applied to the eastern Pacific Bailya: Triton anomalus Hinds, 1844, and Fusus bellus Adams, 1852, the latter placed in synonymy of B. anomala by Keen (1971) and Abbott (1974). [{Fusus bellus Adams, 1852, itself is a junior primary homonym of Fusus bellus Conrad, 1833, an Eocene fossil from Alabama, USA, now referred to Penion (Palmer & Brann, 1966; Synder, 2003)]. But Keen’s (1958, 1971) treatments of the species are confusing. In 1958 she recognized Bailya anomala, with no mention of Fusus bellus, and gave its range as Nicaragua to Panama. She chose to illustrate Hinds’ poor, minute figure (1884a: pl. 4, fig. 14) in both éditions. By 1971 she had included F. bellus as a synonym, perhaps after seeing Turner’ s (1956) illustration of the type of that species, which was based on a juvénile specimen. Keen then changed the range to “Guaymas, Mexico, to Nicaragua” despite the fact that both T. anomalus and F. bellus hâve Panama as their type locality. Although F. bellus is unavailable, the question is which, if either, of the two taxa recognized here is Hinds’ T. anomalus.

The two species presented here are best described as the banded species (Figs 3-8) and the non-banded species (Figs 9 15). Hinds’ figure oï Triton anomalus in the Sulphur expédition account (1844a) is clearly the banded species (Fig 1). Fusus bellus Adams, 1852 (Fig 2), although based on an immature specimen, is also the banded species and thus a junior synonym of Hinds’ taxon as Keen suspected.

A problem arises considering the date and place of publication of Triton anomalus. The description appears in two places, both in July, 1844: in the Sulphur expédition account and in the Proceedings of the Zoological Society of London (PZSL). In Hinds’ description of the species in the Sulphur expédition he lists it as ''Triton anomalus Hinds, /.c.” with the loc. cit. refeiTing to “Proceed. Zool. Soc. Feb. 27, 1844.”

But this refers only to the date his paper was read before the Zoological Society. The actual date of publication was July, 1844, as shown by Duncan (1937), who outlined the tortuous history of the PZSF publication dates. The first part of the Sulphur expédition account was also published in July, 1844. The exact day in July of either publication is unknown. Reeve figured the banded taxon the next month (August, 1844) citing only the Sulphur account. Which came first - the PZSL description or the Sulphur description? Keen (1966) gave the PZSL paper priority, recognizing that the PZSL description dated from July and not February, but without any explanation or proof, perhaps relying on Hinds’ loc. cit. However, because she did not recognize that two species may hâve been involved and makes no point of choosing one date over the other, I do not consider this a nomenclatorial act; in fact, she did not include either reference in her literature cited for that paper.

The identity of the Sulphur account species is obvions because of the figure - it is the banded taxon. But the PZSL account was unfigured. Furthermore, the brief Latin description, although identical in both accounts, seems to apply more to the unbanded species than to the banded one. There is no mention of the prominent bands in the description, which only refers to the shell as "fusca'’’ (brown). But the banded taxon is not brown, although the unbanded one is. Hinds compares his species with Triton fictilis Hinds, 1844 (“its appearance is veiy similar”), a cancellariid that typically does not hâve bands. Thus it is possible that the description does not refer to the banded taxon that Hinds had nonetheless illustrated.

The type of Triton anomalus Hinds, 1844, supposedly at the Natural History Muséum in London, has not been located. Keen (1966) spent six months at the Muséum in 1964-65 looking for Hinds’ material. She was not able to find a significant number of types and remarked that “many of the types and figured specimens were among the lots sold to favored collectors.” But she also noted “I may well hâve overlooked some lots.” After inquiring it was confirmed that the type has not been located (A.

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G.T. Watters

Identity of Triton anomcilus Hinds, 1 844

Salvador, NHMUK, pers. comm., February 2012). The whereaboLits of the type specimen(s) remains Linknown.

Two species of Bailya occur in the eastern Pacific, but which one is Triton anornalusl The identity of the PZSL species is in doubt, whereas the identity of the Siilphur species is not. The dates of the two descriptions cannot be precisely determined. To my knowledge, no previous author has definitively and puiposefully chosen one publication, in terms of precedence, over the other as a nomenclatorial act. In the interest of fixing the identity of Triton anomalus to a spécifie species, I, as First Reviser, déclaré the Sulphur description (1844a: 12) to be the earlier of the two descriptions and designate Hinds’ plate 4, figures 13 and 14 (1844a) as the lectotype of Triton anomalus Hinds, 1844 (ICZN 24.2.2, 73.1.4). Keen (1966) stated that ail of Hinds’ material should be regarded as syntypes; for this reason I hâve designated the figure as a lectotype rather than a holotype. This act also fixes the type species of Bailya, which is Triton anomalus by original désignation of Smith (1944). This act also préservés the identity of Bailya anomala as the taxon to which it has always been associated in the literature and in collections. Triton anomalus is redescribed and the second (unbanded) species is described as new.

Unanswered is the question of the type locality of Triton anomala. If indeed two species were included in the original description, to which species does “Island of Quibo” (= Isla Coiba) pertain? Because of the lack of type material there appears to be no way to solve this problem. Because Isla Coiba lies within the range of both species, I hâve assumed that the type locality of what is here recognized as B. anomala is correct as originally stated.

Abbreviations

GTW: Collection of G. Thomas Watters.

LACM: Natural History Muséum of Los Angeles County, California, USA.

NHMUK: Natural History Muséum, London, UK.

UF: Florida Muséum of Natural History, Gainesville, Florida, USA.

SYSTEMATICS

Family BUCCINIDAE Rafinesque, 1815

Genus Bailya M. Smith, 1944

Subgenus Bailya M. Smith, 1944

Type species by original désignation: Triton anomalus

Hinds, 1844, Recent, eastern Pacific Océan.

Bailya {Bailya) anotnala (Hinds, 1844)

Figs 1-8

Triton anomalus Hinds, 1844a: 12, pl. 4, ligs. 13, 14 (July); Hinds, 1844b: 22 [July]; Reeve, 1844: [83-84],

pl. 20, fig. 100; Reeve, 1847: 97; Smith, 1944: 78; Keen, 1966: 270.

Fusus hellus Adams, 1852a: 353, 531; Adams, 1852b: 129, 307; Adams & Adams, 1853: 78; Carpenter, 1864: 347; Carpenter, 1872: 183; Turner, 1956: 34- 35, pl. 8, fig. 2; Keen, 1971: 557, fig. 1098 [in synonymy of Triton anomalus Hinds, 1844]; Abbott, 1974: 217 [in synonymy of Triton anomalus Hinds, 1844] {non Fusus bellus Conrad, 1833].

Bailya anomala (Hinds, 1844) - Keen, 1958: 398, fig. 530; Keen, 1971: 557, fig. 1098; Abbott, 1974: 217; Watters, 2007: 10, figs. 8,9; Watters, 2009: 225.

Type material. Triton anomalus: Lectotype, designated herein, Hinds, 1844a: pl. 4, figs. 13, 14. Fusus bellus: Holotype, Muséum of Comparative Zoology 177176, illustrated in Turner (1956).

Type locality. Triton anomalus: Island of Quibo [Coiba], VeragLia [Panama]; on the sandy shore at low water.

Fusus bellus: Panama [Pacific].

Distribution and habitat. Eastern Pacific Océan, from at least Guaymas, México, to Ecuador. General ly in shallow water to 30 m, often under dead coral slabs.

Redescription. Average 15.9 mm in length (min, 13.3; max, 19.3). Fusiform; spire ca. 50% total length. Protoconch blunt, white, of 1.5 smooth, slightly shouldered whorls. Teleoconch of 5.5-6 whorls, abruptly arising from protoconch. Teleoconch sculpture of widely separated, spiral cords; 17 1° (primary) cords on final whorl; cords most pro minent on siphonal canal. Between 1° cords is single (occasionally two) 2° (secondary) cords or threads. Both types of cords are more similar in size on the siphonal canal and below the suture. Axial ribs widely spaced; 13-15 ribs on penultimate whorl, 12-14 ribs on last whorl. Axial ribs separated by microscopie axial threads. Sculpture predominately spiral, weakly cancellate, with intersections of spiral and axial sculpture forming slightly more pronounced spiral nodes. Terminal varix well-developed, set back a short distance from outer lip. Aperture oval, weakly or obsoletely crenulated (ca. 6-7 denticles) on outer lip; anal canal set off by two very weak denticles. Columella continuons, smooth. Pariétal callus adhèrent to body whorl for its length. Siphonal canal short, open. Color white beneath a thin, tan periostracLim (usually lost), with 3 dark brown spiral bands, often reduced to spots at the sculptural intersections; individual tubercles on the siphonal canal may be colored as well. Aperture white. Operculum, radula, and anatomy unknown.

Remarks. This species is rather constant in its sculpture and coloration but varies considerably in size. See Table 1 for comparison with B. cidaris. Bailya anomala does not seem to be closely related to

12

G.T. Watters

Novapex 1 4( I ): 11-15,10 mars 20 1 3

any other member of the genus. Its large size and conspicLioLis color bands are unique.

Bailya (Bailya) cidaris n. sp.

Figs 9- 15

Type niaterial. Holotype LACM 3243, 3 m, Isla Coiba, Veraguas Provinee, Panama, 3 m.

Paratypes: 1 LACM 3244, 20-30 m, Bahia de Chiriqui, Panama, 15.8 mm length; 1 OSUM 37272, 20-30 m, Bahia de Chiriquf, Panama, 15.3 mm length; 1 UF 451539, 20-30 m, Bahia de Chiriqui, Panama, 14.7 mm length; 1 OSUM 37273, 6-7 m, off Puntarenas Province, Costa Rica, 13.7 mm length.

Type locality. Eastern Pacific Océan, Isla Coiba, Veraguas Province, Panama, 3 m.

Distribution and habitat. Eastern Pacific Océan. Knovvn only from Bahia de Chiriqui and Isla Coiba, Panama, to Costa Rica. Specimens hâve been found from 3 to 30 m; live specimens were collected in 3 m. Based upon the type locality of B. anomala, both species occur at Isla Coiba.

Description. Average 14.9 mm in length (min, 13.7; max, 15.8; holotype 15.2 x 7.0 mm). Fusitorm; spire ca. 50% total length. Protoconch blunt, white or tan, of 1 .5 smooth, slightly shouldered vvhorls. Teleoconch of 6 whorls, abruptly arising from protoconch. Teleoconch sculpture of widely separated, raised, spiral cords; 11 12 1° cords on final whorl; cords most prominent on siphonal canal. Between 1° cords are 3 more or less equally sized 2° cords or threads. Axial ribs widely spaced; 13-14 ribs on penultimate whorl, 12-13 ribs on last whorl. Axial ribs separated by microscopie axial threads. Sculpture cancellate, with intersections of spiral and axial sculpture forming prominent tubercles. Terminal varix well-developed, set back a short distance from outer lip. Aperture oval, weakly crenulated (ca. 6 denticles) on outer lip; anal canal set off by two very weak denticles. Columella continuons, smooth. Pariétal callus adhèrent to body whorl for its length. Siphonal canal short, open. Color tan, axial ribs often dark brown, with or without 4 darker spiral bands, spiral cords and tubercles may be lighter colored. Aperture white. Operculum tan to dark red, leaf-shaped, with terminal apex. Radula and anatomy unknown.

Figures 1-15

1-8. Bailya anomala (Hinds, 1 844).

1. Figures 13, 14 of Triton anomalus from Hinds, 1844a. 2. Holotype MCZ 177176 of Fnsns hellus Adams, 1852. Panama, ca. 11 mm length, subadult, figure from Turner (1956). 3. GTW 9114a, 18 m, Isla Venado, Panama, 12.5 mm length, subadult. 4, 5. GTW 9114b, 1-2 m, Puerto Vallarta, México, 19.3 mm length. 6. GTW 9114c, 1 m, Isla Pedro Gonzales, Panama, 16.6 mm length (with periostracum). 7. GTW 91 14e, Bahia Rincon, Costa Rica, 14.5 mm length. 8, Detail of siphonal canal sculpture from GTW 9114b.

9-15. Bailya cidaris n. sp.

9, 10. Holotype, LACM 3243, 3 m, Isla Coiba, Panama, 15.3 m length. 11, 12. Paratype, LACM 3244, 20-30 m, Bahia de Chiriquf, Panama, 15.8 mm length. 13. Paratype, OSUM 37272, 20-30 m, Bahia de Chiriquf, Panama, 15.3 mm length. 14. Paratype, UF 451539, 20-30 m, Bahia de Chiriquf, Panama, 14.7 mm length. 15. Detail of siphonal canal sculpture from LACM 3244.

13

G.T.Watters

Identity of Triton anomalus Hinds, 1844

Remarks. Bailya cidaris differs from the only other known Bailya in the eastern Pacific Océan {B. anomala) in color and sculptural details (Table 1). It appears to hâve a much more restricted distribution than B. anomala, being limited to the coast of Panama and Costa Rica. It’s closest relative, based on shell features, is the western Atlantic Océan Bailya inîricata (Dali, 1884), particularly specimens from eastern Panama (see Watters, 2009: figs. 131-132).

Bailya intricata differs from B. cidaris in having more 1° spiral cords on the last whorl (12-16 vs 11-12) and more axial ribs on the last whorl (13-16 vs 12-13). The axial ribs in B. intricata are rarely darkly colored as in 5. cidaris.

Etymology. L. cidaris, a Persian diadem or tiara. A féminine noun in apposition.

Table 1. Shell différences between B. anomala and B. cidaris. Spiral sculpture - cords arranged by decreasing size A to B.

Shell characteristic	Bailya anomala	Bailya cidaris
Length (mm)	13.3-19.3	13.7-15.8
Number of axial ribs on final whorl	12-14	12-13
Number of axial ribs on penultimate whorl	13-15	13-14
Spiral sculpture pattern on body whorl	A-B-A	A-B-B-B-A
Intersections form prominent tubercles	No	Yes
Sculpture cancellate	Weak	Strong
Number of 1° spiral cords on final whorl	17	11-12
Color pattern	White with three broad interrupted brown spiral bands	Tan with narrow pale bands, axial ribs may be darker

Acknowledgments

1 am indebted to Ms. K. Way and Ms. A. Salvador, NHMUK, for their attempts to locate the type of Triton anomalus, L. Groves (LACM) and J. Slapcinsky (UF) for catalog numbers, and particularly to R. Petit, North Myrtle Beach, SC, USA for invaluable guidance in preparing this paper. The author is grateful to the two reviewers, James H. McLean and Koen Fraussen, for valuable improvements to the manuscript.

REFERENCES

Abbott, R.T. 1974. American Seashells. 2"^ ed. Van Nostrand, New York: 663 pp.

Adams, C. B. 1852a. Catalogue of shells collected at Panama, with notes on synonymy, station, and habitat. Annals of the Lyceiim of Natural History of New York 5: 229-296 [lune], 297-549 [July]. Adams, C. B. 1852b. Catalogue of Shells Collected at Panama, with Notes on Synonymy, Station, and Geographical Distribution. R. Craighead, New York: viii-i-334+errata.

Adams, H. & Adams, A. 1853. The Généra of Recent Mollusca, Arranged According to Their Organiz.ation. London. 1 . Part 111: 65-96, pis. 9-12. Carpenter, P. P. 1864. Review of Prof. C. B. Adam’s

“Catalogue of the shells of Panama,” from type specimens. Proceedings ofthe Zoological Society of London for 1863: 339-369.

Carpenter, P. P. 1872. The mollusks of western North America. Embracing the second report made to the British Association on this subject, with other papers; reprinted by permission, with a general index. Smithsonian Institute Miscellaneous Collections 10(252): xii+325-f-13-121 .

Conrad, T. A. 1833. Fossil Shells ofthe Tertiary Formations of North America. Philadelphia. 1(4): 39-46.

Duncan, F. M. 1937. On the dates of publication of the Society’s 'Proceedings,'’ 1859-1926. With an appendix containing the dates of publication of 'Proceedings,’’ 1830-1858, compiled by the late F. H. Waterhouse, and of the 'Transactions,’ 1833- 1869, by the late Flenry Peavot, originally published in P. Z. S. 1893, 1913. Proceedings of the Zoological Society of London 107: 71-84.

Hinds, R. B. 1844a. The Zoology ofthe Voyage ofthe H.M.S. Sulphur, Under the Command ofCapt. Sir Edward Belcher, R.N., C.B., F.R.G.S., etc. During the Years 1836-1842. Mollusca. London. Part 1: 1- 24, pis. 1-7 [July], Part 11: 25-48, pis. 8-14 [Oct.].

Hinds, R. B. 1844b. Six species of Triton, Solarium, and Corbula. Proceedings ofthe Zoological Society of London, part Xll: 21-26.

14

G. T. Watters

Novapex 14(1): 1 1-15, 10 mars 2013

Keen, A. M. 1958. Sea Shells of Tropical West America. T' ed. Stanford University Press, Stanford: 624 pp.

Keen, A. M. 1966. West Ameriean mollusk types in the British Muséum (Natural History) 11. Species described by R. B. Hinds. Veliger 8: 265-275, 5 figs., pis. 46, 47.

Keen, A. M. 1971 . Sea Shells of Tropical West America. 2™' ed. Stanford University Press, Stanford: 1064 pp.

Palmer, K.V. & Brann, D.C. 1966. Catalogue of the Paleocene and Eoeene mollusks of the Southern and eastern United States. Part IL Gastropoda. Bulletin of American Paleontology 4?>: 471-1057.

Reeve, L. A. 1844. Monograph of the genus Triton. Conchologica Iconica 2: 45-84, index, pis. 1-20.

Reeve, L. A. 1847. Eléments of Conchology . London. 1. Part 7: 97-112.

Smith, M. 1944. Panamic Marine Shells. Synonymy, Nomenclature, Range and Illustrations. Tropical

Photographie Laboratory, Winter Park, Llorida: xiii-i-127 pp., 912 figs.

Snyder, M. A. 2003. Catalogue of the marine gastropod family Lasciolariidae. Academy of Natural Sciences, Spécial Publication 21 : 431 pp.

Turner, R. D. 1956. The eastern Pacific marine mollusks described by C. B. Adams. Occasional Papers on Mollusks 2(20): 21-1 35, pis. 5-21 .

Watters, G.T. 2007. Small western Atlantic

Buccinidae. Part 1 . The genus Bailya M. Smith, 1944. American Conchologist 35(3): 10-1 1, 12 figs.

Watters, G.T. 2009. A révision of the western Atlantic Océan généra Anna, Antillophos, Bailya, Caducifer, Monostioliim, and Parviphos, with description of a new genus, Dianthiphos, and notes on Engina and Hesperisternia (Gastropoda: Buccinidae: Pisaniinae) and Cumia (Colubrariidae). A^«wr/7w.s' 123: 225-275.

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E. Rolân, R. Fernàndez-Garcés & F. Rubio

Novapex 14(1): 17-19, 10 mars 20 13

A new species of Murchisonella (Heterobranchia, Murchisonellidae)

from Cuba

Emilio ROLÂN Museo de Historia Natural,

Parque Vista Alegre, Universidad Campus Norte,

15782 Santiago de Compostela, Spain

Raül FERNANDEZ-GARCÉS Centro de Estudios Ambientales de Cienfuegos (CEAC),

Division de Gestion Ambiental (DGA), calle 17, esquina Ave. 46,

Cienfuegos, Cuba

Federico RUBIO Pintor Ribera,4, 16a 46930 Quart de Poblet, Valencia, Spain

Keywords. Heterobranchia, Murchisonellidae, Murchisonella, new species, Cuba.

Abstract. A new species of the genus Murchisonella is described and compared with the other species known from the Caribbean.

Resumen. Se describe una nueva especie del género Murchisonella comparândola con la otra especie ya conocida del Caribe.

INTRODUCTION

Recently a world-wide révision of the genus Murchisonella (Peiïas & Rolân, in press) showed that only one species is known from the Caribbean waters: Murchisonella specîrum Môrcb, 1875 with the synonyms Aclis bermudensis Dali & Bartsch, 1911 and Bermudaclis tarnpaensis Bartsch, 1947. This species is well known and presented in many publications (for example, Redfern, 2001 : 151,pl.67). In the collection of samples taken by the second author (RFC) from several localities in Cuba some shells of an unknown species of this genus were found. The préservation of these shells indicates they may hâve originated from quaternary deposits.

Abbreviations

MCZ: Muséum of Comparative Zoology, Harvard University, Cambridge, U.S.A.

MHNS; Museo de Historia Natural, Santiago de Compostela, Spain.

MNCN: Museo Nacional de Ciencias Naturales, Madrid, Spain.

MNHN; Muséum National d'Histoire Naturelle, Paris, France.

IFS: InstitLito de Ecologia y Sistemâtica, Havana, Cuba.

USNM: National Muséum of Natural History, Washington, D.C., U.S.A.

CFG: collection of R. Femândez-Garcés. s: empty shell.

SYSTEMATICS

Family MURCHISONELLIDAE Casey, 1904

Genus Murchisonella Môrch, 1875

Type species: Aclis {Murchisonella) spectruni Môrch. 1875, by monotypy.

Murchisonella ultiina spec. nov.

Figs 1-4

Type material. Holotype (Fig. 1) in MNCN (15.05/60062, s). Paratypes: MNHN (IM-20 12-28, 1 s, Fig. 2); MHNS (100576, 3 s); CFR (3 s); ail from the type locality. Other paratypes: IFS (1 s) CFG (1 s) from Calicito, Cienfuegos Bay, 22°07.970’N, 80°29.824’W, 12 m, CubÂ; MCZ (1 s) from Los Pinitos, Cienfuegos Bay, 22°07.886’N, 80°27.313’W, 8-10 m, Cuba; CFG (5 s) Cienfuegos Bay, Cuba.

Type locality. North of Cuba, mouth of the Sagua River, Villa Clara Province, 22°57’36”N, 80°00’47”W, in sédiments 4.5 m depth.

Etymology. The species name dérivés from the Latin ultinms, -a, -uni, alluding to be the last species found after a large révision of this group (in press).

17

E. Rolàn, R. Fernândez-Garcés & F. Rubio

A new species of Miirchisonella from Cuba

Description. Shell (Figs 1-3) small, subcylindrical, whitish and fragile. Protoconch (Fig. 4) typical for the genus, smooth, at one level, with a diameter of about 200 ;<m, small nucléus (diameter about 20 //m) and a clear séparation from the teleoconch. Teleoconch with about 4 yi- 5 whorls, slightly convex, vvhorl height slowly increasing, from the start having about 9 spiral grooves that are regularly distributed over the whorl and are crossed by numerous and irregular axial folds. From the second whorl the folds on the upper part of the whorls (between the 4 upper grooves) are clearly opistocyrt (curved backward), while those of the lower part are prosocyrt (curved forward). These folds are narrow and numerous (about 130 by whorl). There is no clear séparation between the upper and lower parts of the whorls except for the switch in microsculpture curvature. Aperture rounded; columella slightly curved lacking any fold or tooth.

Dimensions: The holotype is 1.25 mm high. The other shells hâve a similar size.

Distribution. Only known from the type material, from the north and south coasts of Cuba.

Remarks. The only known congeneric species in the Caribbean (M. spectrum) is larger (up to 3 mm), with a more conic profile, an évident step separating the upper and lower parts of the whorls, less and wider spiral grooves, and less developed axial sculpture.

We wanted to make a comparison with Henrya morrisoni Bartsch, 1947 (Murchisonellidae) which has a similar size, but the holotype of that species in the USNM (sample 573636) has been lost (pers. comm. Ellen Strong and Yolanda Villacampa). In the description of this species Bartsch (1947) mentions that its whorls are “strongly rounded” and the “suture

strongly constricted”, which differentiates it from our new species. Shells of 1.4 mm high hâve 5.4 whorls. The specimen of H. morrisoni illustrated in Redfern (2001: plate 67) has strongly convex whorls and constricted suture and therefore probably is Henrya henryi. In the original description of H. morrisoni there is no mention of spiral grooves as are présent in the species here described. The two other species of this Caribbean genus described by Bartsch (1947) are H. henryi and H. goldmani. Both are larger (the types measure 2.1 mm high with 6.3 whorls {B. henryi) and 2.0 mm high with 6 whorls {H. goldmani) and the whorls are strongly convex.

Acknowledgements

Jésus Méndez of the CACTI (Centre de Apoyo CientiTico y Tecnologico a la Investigaciôn, Universidad de Vigo) made the SEM photographs. We also thank Han Raven (Naturalis Biodiversity Center, Leiden, The Netherlands) for his help in the translation and for other useful remarks.

REFERENCES

Bartsch, P. 1947. A monograph of the West Atlantic Mollusks of the family Aclididae. Smithsonian Miscellaneous Collections , 106(20): 1-29, 5 pis. Penas, A. & Rolân, E. (in press). Révision of the subfamily Murchisonellinae Casey, 1904 (Mollusca, Gastropoda, Heterobranchia, Murchisonellidae). Vita Malacologica.

Redfern, C., 2001 . Bahamian Seashells. A thousand species from Abaco, Bahamas. Bahamian seashells, Boca Raton, 280 pp, 124 pis.

Figures. 1-7. Murchisonella iiltiina spec. nov.

1. Holotype, 1 .25 mm, Sagua (MNCN); 2. paratype, 1 .2 mm, Sagua (MNHN); 3. paratype, 1 .3 mm, Calicito, Cienfuegos Bay (IFS); 4. apex and protoconch; 5-7. microsculpture and detail.

18

No V APEX 14(1): 17-19, 10 mars 20 13

E. Rolân, R. Fernândez-Garcés & F. RUBIO

19

<(•

J. Phlorce

Novaf’EX 14(1): 21-24, 10 mars 2013

Deux nouvelles espèces de Columbellidae (Gastropoda: Neogastropoda)

de l’île de Martinique, Antilles françaises

Jacques Pelorce

289, voie les Magnolias 3024Ü Le Grau du Roi pelorce@free.fr

MOTS CLÉS. Gastropoda, Neogastropoda, Columbellidae, Martinique, Costoanachis n.sp., Astyris n.sp.

RÉSUMÉ. Deux nouvelles espèces de Columbellidae du genre Anacliis sous genre Costoanachis et du genre Astyuis sont décrites de l’île de Martinique, Antilles françaises.

ABSTRACT. Two new species of Columbellidae of the genus Anachis subgenus Costoanachis and genus Astyris are described from Martinique Island, French Antillean.

INTRODUCTION

En 2006, Régis Delannoye, pêcheur martiniquais, remonte de façon accidentelle de vieux filets perdus par 240 m de profondeur au large de La Trinité côte nord-est de l’île de la Martinique. A l’examen de ces vieux filets concrétionnés, de nombreux spécimens de diverses espèces de mollusques furent récoltés. Parmi ceux-ci figuraient une vingtaine de spécimens, de taille inférieure au centimètre, appartenant à deux espèces inconnues de la famille des Columbellidae.

Abréviations

CJP: collection Jacques Pelorce

CKM; collection Kevin Monsecour

MNFTN: Muséum national d’Histoire naturelle, Paris,

France

SYSTEMATIQUE

Famille COLUMBELLIDAE Swainson, 1840 Sous-famille ATILIINAE Cossmann, 1901 Genre Anachis H. & A. Adams, 1853 Sous-genre Costoanachis Sacco, 1 890 Espèce type par désignation subséquente Columbella (Anachis) turrita Sacco, 1890 (Pace, 1902: 43), non Sowerby, 1832 (= Costoanachis saccostata nom. nov) (Radvvin, 1968 : 120) fossile du tertiaire du Piémont et de la Ligurie (Italie).

Discussion. Le sous-genre Costoanachis introduit par Sacco (1890) de manière très succincte « Anfractus omnes longitudinal iter costati excepta C. corrugata Brocchi var. B)» a été redéfini par Radvvin (1968: 120): "Shell smal! to moderate in size (4- 15mm in length), and convex, suture shallovv to incised or impressed; body vvhorl equal to or less than Vi total length, aperture moderately wide; columella straight, weakly denticulate, siphonal canal short and straight; sculpture of prominent axial ribs, in some cases limited to the body vvhorl, commonly vvith

microscopie spiral grooves between them. Color variable". Nos spécimens correspondent parfaitement à cette description sauf en ce qui concerne la hauteur du dernier tour qui est supérieure à la moitié de la hauteur totale. Cependant dans sa définition du genre Radvvin (1968: 120) inclut et figure dans ce genre les espèces C. avara (Say, 1822), C. hotessieriana (Orbighy, 1842), C. sparsa (Reeve, 1859) ou C. sciitulata (Reeve, 1859), entre autres, qui ont les derniers tours dont la hauteur est supérieure à 60 % de la hauteur totale.

Anachis (Costoanachis) martinicensis n. sp.

Figs 1-8

Localité type. Au large de La Trinité, côte nord est de l’île de la Martinique, 240 m de profondeur.

Matériel type. Holotype (6,4 x 3,0 mm) MNHN 25212 (Figs 1-5), paratype 1 (6,1 x 2,7 mm) (Fig. 6) CJP, paratype 2 (7,0 x 3,0 mm) (Fig. 7) CJP, paratype 3 CJP, paratype 4 MNHN et paratype 5 CKM, tous de la localité type.

Distribution, Connue actuellement uniquement de la localité type au large de La Trinité, au nord-est de l’île de la Martinique, par 240 m de profondeur.

Description. Coquille de petite taille (L = 6.0 à 7.1 mm), largeur sensiblement égale à 44 % de la hauteur, dernier tour occupant environ 60 % de la hauteur totale, suture incisée bien marquée, tours légèrement convexes à la base et régulièrement étagés. Protoconque bulbeuse, composée d'environ 1 tour et demi, blanche à beige clair, surface lisse. Sculpture de la téléoconque constituée de très fines stries d’accroissement, de 32 stries spirales, 12 sur l’avant dernier tour, les 4 premières sous la suture plus serrées et une vingtaine sur la moitié inférieure du dernier tour, de 10 grosses côtes axiales aiTondies sur le premier tour puis 7 sur les tours suivants où elles sont alignées, l’espace entre les côtes est égal au double

21

J . Pelorce

Deux nouvelles espèces de Columbellidae de Martinique

environ de la largeur de celles-ci. Ouverture ovale, canal anal légèrement marqué, canal siphonal court et profond. Sur les spécimens matures le labre épaissi extérieurement porte à l’intérieur 8 plis très légers, un cal columellaire débordant légèrement l’ouverture, un pli peu marqué à la base de la columelle. Couleur blanc à beige clair avec quelques taches irrégulières marron clair.

Discussion. Une seule espèce récemment décrite de la Guadeloupe ressemble à la nouvelle espèce : C. roberti Monsecour & Monsecour, 2006, C. martinicensis s’en différencie aisément par sa taille plus petite, un nombre de côte plus réduit, des stries spirales moins apparentes, des tours moins convexes et une coloration moins vive.

Etymologie. L’espèce est nommée en référence à l’île de la Martinique d’où proviennent les spécimens.

Genre Astyris H. & A. Adams, 1853 Espèce type par désignation subséquente Astyris rosacea (Gould, 1840) (Cossmann, 1901: 238), Groenland au New Jersey; Alaska; Nord de l'Europe.

Astyris delannoyei n. sp.

Eigs 8-14

Localité type. Au large de La Trinité côte nord est de la Martinique, 240 m de profondeur

Matériel type. Holotype (8,0 x 3,5 mm) MNHN 25213 (Figs 8-11), paratype 1 (7,4 x 3,4 mm) (Fig. 12) CJP, paratype 2 (6,7 x 3,1 mm) (Fig. 13) CIP, paratype 3 (6,0 x 2,8 mm) (Fig. 14) CJP, paratype 4 MNHN, paratype 5 CKM, paratypes 6 à 12 CJP, tous de la localité type.

Distribution. Connue actuellement uniquement de la localité type au nord-est de l’île de la Martinique par 240 m de profondeur

Description. Coquille de petite taille (L = 6.0 à 8.0 mm), largeur sensiblement égale à 45 % de la hauteur.

dernier tour occupant environ 2/3 de la hauteur totale, suture incisée bien marquée, tours légèrement convexes et régulièrement étagés, coquille brillante, translucide; fausse suture nettement visible par transparence sur les premiers tours, suture incisée bien marquée. Protoconque élevée, composée d'environ 3 tours, blanc crème, surface lisse. Sculpture de la téléoconque constituée de très fines stries d’accroissement et d’environ 10 à 14 stries spirales sur la moitié inférieure du dernier tour. Ouverture ovale, canal anal très légèrement marqué, canal siphonal court et profond. Sur les spécimens matures le labre épaissi extérieurement porte 2 à 4 dents à l’intérieur de la partie supérieure, cal columellaire appliqué, deux plis bien marqués à l’intérieur de l’ouverture à la base de celle-ci. Décoration très variable composée de taches et de lignes imégulières marron et de taches blanches sur fond beige, une ligne marron plus foncée au dessus de la suture peut être présente.

Discussion. Deux espèces de la région caraïbe ressemblent à la nouvelle espèce: Astyris lunata (Say, 1826) (Figs 15-16) et A. multilineata (Dali, 1889) (Figs 17-18). Ces deux espèces sont figurées et décrites par Tunnel et al. (2010: 216-217). Elles se distinguent d’A. dellanoyei par leur taille beaucoup plus petite, par leur protoconque qui possède un tour de moins, et par une suture des tours de téléoconque non incisée.

Etymologie. L’espèce est nommée en l’honneur de Régis Delannoye, pêcheur martiniquais de la Trinité, qui a remonté le filet perdu dans lequel il a trouvé les spécimens décrits ici.

Remerciements

Je voudrais remercier Franck Boyer qui m’a transmis les spécimens récoltés par Régis Delannoye, le professeur Philippe Bouchet et Virginie Héros du Muséum national d’Histoire naturelle de Paris pour leur aide et leurs encouragements, le référé Kevin Monsecour pour ses remarques sur le manuscrit et enfin Roland Houart pour son aide éditoriale.

Figures 1-18

1-7. Anachis (Costoanachis) martinicensis n. sp. Nord-Est Martinique, 240m.

1-5. Holotype MNHN 25212, 6,4 x 3,0 mm; 6. Paratype 1 CJP 6,1 x 2,7 mm. 7. Paratype 2 CJP 7,0 x 3,0 mm. 8-14. Astyris delannoyei n. sp. Nord-Est Martinique, 240m.

8-11. Holotype MNHN 25213, 8,0 x 3,5 mm; 12. Paratype 1 CJP 7,4 x 3,4 mm. 13. Paratype 2 C.TP 6,7 x 3,1 mm. 14. Paratype 3 CJP 6,0 x 2,8 mm.

15-16. Astyris lunata (Say, 1826), St Kitts CJP 3,3 x 1 ,65 mm.

17-18. Astyris multilineata (Dali, 1 889), Antigua CJP 3,5 x 1 ,45 mm.

22

J. Pelorce

Novapex 14(1); 21-24, 10 mars 20 13

••

J. PCLORCE

Deux nouvelles espèces de Columbellidae de Martinique

REFERENCES

Cossmann, M. 1901 . Essais de Paléontologie comparée Quatrième livraison: 221-241 .

Pace S. 1902 Contributions to the Study of the Columbellidae N°1 Proceedings ofthe Malacological Society’ Vol. V Part 1 : 36-1 1 1

Radwin G.E. 1968 New Taxa of Western Atlantic Columbellidae (Gastropoda, Prosobranchia) Proceedings ofthe Biological society of Washington Vol. 81 : 143-150

Sacco F. 1890 (dans Bellardi L.) I Molluschi dei TeiTcni Terziari del Piemonte e délia Liguria Memorie délia Reale Academia delle Scienze di Torino Sérié seconda Tomo XL : 295-368 Tav. I et II

Tunnell J.W., Andrews J., Barrera N.C., Moretzsohn F. 2010. Encyclopedia of Texas Seashells. Texas A&M University Press. 1:512 pp.

24

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Keen, A.M. & Campbell, G. B. 1964. Ten new speciesof Typhinae (Gastropoda : Muricidae). TheVeliger7(\): 46-57.

Po\A/ell, A.W.B. 1979. New Zealand Mollusca. Marine, land and freshwater shells. William Collins Publishers Ltd: xiv + 500 pp.

Mayr, E. 1989. Attaching names to objects. In: What the philosophy ofbiology is : essays for David Hull (M. Ruse, ed.), Klumer Academie, Dordrecht: 235-243.

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Papers describing new species (subspecies) will be accepted only if the primary types are deposited in a recognized public Muséum or scientific Institution.

The paper will be in accordance with the ruies of the International Code ofZoological Nomenclature (Fourth édition)

Manuscripts. Manuscripts will be in English or in French. They must be typed on one column, ragged right (left-justified), double-spaced throughout, on one side only of A4. Margins must be at least 25 mm. The sequence of sections will respect the following order: title, name of author(s), address(es) of author(s), keywords and summary in English. Generic and (sub)specific names hâve to be typed in italics. References in the text should be given as follows: Keen & Campbell (1964) or (Keen & Campbell, 1964). Refer to a recent issue of Novapex for the lay out.

References, in alphabetic order, should be given in the following form (titles of journals should not be abbreviated):

Keen, A.M. & Campbell, G. B. 1964. Ten new species of Typhinae (Gastropoda : Muricidae). TheVeliger7{\): 46-57.

Powell, A.W.B. 1979. New Zealand Mollusca. Marine, land and freshwater shells. William Collins Publishers Ltd: xiv + 500 pp.

Mayr, E. 1989. Attaching names to objects. In: What the philosophy of biology is : essays for David Hull (M. Ruse, ed.), Klumer Academie, Dordrecht: 235-243.

Internet resources.

Author(s) 2007. Title of the website or database, publisher name and locatiuon (if indicated), number of pages, address: http://www.... (date of access).

Illustrations. Photographe must be of a high quality (colour or black/white), printed on glossy paper in a final version (max. 16 X 21 cm), adequatly mounted. Colour work can be submitted for black & white production. The illustrations may be submitted as digital files (CD-ROM, ZIP) in BMP, JPG or TIFF format, with mention of the program. They must be adequately mounted with not any other text than the numbering. A printed version of the plates must be imperatively sent together with the manuscript.

If more than one figure is included in a plate or/and in the text, ALL figures must be consecutively numbered (Fig. 1, 2, 3...) NOT Fig IA, IB, IC, NOR plate 1, Fig. 1... Inclusion of colour plates has to be approved by the board who will take the final decision. Authors who want to include colour plates are invited to ask for possibilities and charges.

Processing of manuscripts. Manuscripts will be submitted to the board who will distinguish between the articles of scientific interest, and those of general aim. The commente will be communicated to authors, who will consider them. A diskette containing the corrected version should be sent back to the Belgian Malacological Society (in Word for Windows support) together with a printed copy. It should strictiy follow the style instructions which will be communicated to the author(s).

Reprints. With regard to papers of scientific interest, 30 reprints are free of charge, representing a maximum of 240 pages, if at least one author is member of the Society. Additional copies (at least 30) will be invoiced at cost.

For non-members, the reprints (min. order 30 copies) will be billed to the author(s). Mailing costs are aiways to be paid by authors.

Manuscripts and any mail are to be sent to:

Société Belge de Malacologie, Mr. C. Vilvens, rue de Hermalle, 113, B-4680 Oupeye, Belgium

or by e-mail: vilvens.claude@skynet.be

	NOVAPEX/SOCIETE
	(suite)
M. Alexandre	L'écho des réunions ;
	- Christiane Delongueville et Roland Scaillet ; Richesse	5
	malacologique sur les côtes du détroit de Gibraltar
C. Vilvens &	Morceaux choisis (quand la presse parle de mollusques)	6
E. Meuleman	Sr
C. Vilvens	^ Quoi de neuf ? La bourse d'Antwerpen	7
R.Scaillet	, Quelques nouvelles publications	8
E. Meuleman	Nous avons reçu	9
C. Delongueville &	- ^ Les marées de 20 1 3	16
R. Scaillet	S

Nov APEX / Société 14(1), 10 mars 20 1 3

Prochaines activités de la SBM

Claude VILVENS

Lieu de réunion : A partir de 14h, à notre local habituel :

Salle **MemIing” (1er étage - ascenseur) - Rue de Genève, 470b - Schaerbeek (Bruxelles)

SAMEDI 27 AVRIL 2013

M. Alexandre : Un panorama des Haliotidae.

Notre trésorier s'intéresse à cette famille depuis longtemps. Pour ceux qui n'étudient pas particulièrement les Haliotis, elles se ressemblent toutes. Notre spécialiste nous dévoilera quelques-uns de ses secrets de détermination de ces belles coquilles "aux grandes oreilles" et qui semblent pouvoir "se découper selon les pointillés" formés par leurs petits orifices si parfaitement alignés ;-) !

***

SAMEDI 25 MAI 2013

Tout le monde : L'excursion de printemps de la SBM.

Avec le beau temps revient l'envie d'aller sur le terrain ;-) (air connu !). Plusieurs zones sont envisagées : notre équipe de reconnaissance déterminera en avril celle qui semble la plus intéressante (plus tôt, c'est trop tôt pour les escargots ;-)).

Durée de l'excursion : la matinée (fin vers 12h30).

Pour ceux qui sont intéressés par les mollusques d'eau douce, prévoir des bottes et surtout une épuisette.

Comme d'habitude, les informations fraîches seront disponibles sur notre site Internet (http://www.societe-belge- de-malacologie.be/) ou auprès de Claude (vilvens.claude@skynet.be ou 04/248.32.25) et Etienne Meuleman (e.meuleman@skynet.be ou 04/380.55.16). Comme d'habitude aussi, il convient de prévoir d'emporter sa bonne humeur, un guide de détermination ... et sans doute aussi bottes et vêtements de pluie (en principe, il fera doux et ensoleillé, mais bon ;-) . . .).

SAMEDI 15 JUIN 2011

Tout le monde : Le grand jeu des coquillages

Avant les grandes vacances, nous vous proposons de nous retrouver pour une activité ludique : nous allons en effet jouer au coqui-quiz-lotto - vous l'aurez compris, il s'agira d'un quiz malacologique, formule qui a remporté beaucoup de succès lors de ses précédentes éditions. Et on gagne quelque chose ; mais oui, des coquilles bien sûr © Un joyeux après-midi avant les grandes vacances !

Pour les informations de dernière minute :

À,

m -■;>

Sur Internet

http://www.societe-belge-de-malacologie.be/

2

Novapex / Société 14(1), 10 mars 2013

Novapex/Société : la publication généraliste de la SBM

Rédacteurs en chef : Claude Vilvens & Etienne Meuleman

Tous les articles généraux sont les bienvenus pour Novapex/Société © !

Afin de faciliter le travail de la Rédaction, il est vivement (et le mot est faible ;-)) souhaité de respecter les règles suivantes pour les articles proposés :

♦ document MS-Word (pour PC Windows 2000 ou XP);

♦ police de caractères Times New Roman;

♦ texte de taille 10, titres de taille 12;

♦ interligne simple;

♦ toutes les marges à 2,5 cm;

♦ document en une seule section;

♦ pas de mode colonne;

♦ photos en version électronique JPG.

Merci pour les Scribes ;-) ! N'hésitez pas à demander une page avec en-tête pour cadrer au mieux vos travaux (vilvens.claude@skynet.be ou e.meuleman@skynet.be).

Petit rappel du trésorier

Marc ALEXANDRE

Chers amis,

Pensez à renouveler votre cotisation dès maintenant.

Nous vous rappelons aussi que celle-ci s’est vu augmenter de 3 euros pour 2013 et sera donc de 43 euros pour les membres habitant en Belgique et de 58 euros pour les membres résidant à

l’étranger.

De même, je vous rappelle également notre changement de eompte qui se trouve dorénavant à la

Banque ING, Bruxelles.

Merci d'effectuer vos versements

sur le compte n° 363-0831752-17 [IBAN: BE61 3630 8317 5217 - BIC : (= SWIFT) BBRUBEBB] au nom de M. Marc ALEXANDRE, rue de la Libération, 45, 6182 Souvret Belgique.

Merci de bien vouloir porter une attention toute particulière à ees modifications afin de nous éviter des

désagréments de rappel.

Novapex / Société 14(1), 10 mars 2013

3

Extension de l’aire de distribution de Bornia aartseni Gofas, 2012 aux côtes atlantiques du Sud de la Péninsule Ibérique

Christiane DELONGUEVILLE

Avenue Den Doom, 5 - B - 1180 Bruxelles - christiane.delongueville@skvnet.be

Roland SCAILLET

Avenue Franz Guillaume, 63 - B - 1140 Bmxelles scaillet.roland@,skvnet.be

MOTS CLEFS/ KEY WORDS

Bornia aartseni - Kelliidae - Algarve - Portugal

RÉSUMÉ

Un exemplaire intact et 4 valves de Bornia aartseni (Bivalvia - Kelliidae) ont été récoltés sur la plage de Monte Gordo en Algarve (Portugal). L’aire de distribution de Bornia aartseni, précédemment limitée à l’Andalousie méditerranéenne, est élargie à la côte atlantique du sud de la Péninsule Ibérique.

ABSTRACT

One intact specimen and 4 valves of Bornia aartseni (Bivalvia - Kelliidae) were found on the beach of Monte Gordo, Algarve (Portugal). The distribution range of Bornia aartseni, previously limited to the Mediterranean coast of Andalusia now extends to the Atlantic coast of the Southern Iberian Peninsula.

INTRODUCTION

La famille des Kelliidae (Veneroida - Galeommatoidea) qui regroupe de petits bivalves a été largement illustrée par van Aartsen en 1996 et par Gofas en 201 1. En Europe, quatre genres composent la famille ; Kellia Turton, 1822, Pseudopythina P. Fischer, 1878, Bornia Philippi, 1836 et Solecardia Conrad, 1849.

Dans les eaux du nord-est Atlantique et de Méditerranée, le genre Bornia Philippi, 1836 est représenté par quatre espèces (CLEMAM consultation 01-2013) :

- Bornia sebetia (Costa O. G., 1829): espèce de forme triangulaire quasi équilatérale, à la coquille brillante, présente en Algarve, sur la côte atlantique du Maroc et en Méditerranée.

- Bornia geoffroyi (Payraudeau, 1 826) : espèce de forme plus arrondie que la précédente, avec umbo prosogyre, présente en Atlantique, de l’Algarve au Sénégal.

- Bornia canariensis D.F. Eloeksema et Simons, 2011 : espèce de forme assez similaire à Bornia sebetia, présente dans l’archipel des Canaries,

- Bornia aartseni Gofas, 2012 ; espèce de Méditerranée récemment décrite, présente en Andalousie (Benalmâdena et environs).

Bornia aartseni se différencie de Bornia sebetia par les caractères suivants (Gofas 2012) :

- une taille plus petite (3 mm versus 8 mm),

- une prodissoconque plus petite (260-280 pm versus 450-500 /ym),

- une ligne palléale proportionnellement plus proche du bord ventral,

- une surface interne des valves quelque peu givrée et dépourvue de fines lignes radiaires,

- une épaisseur et une orientation différentes des dents dans la charnière. Dans la valve gauche, une caractéristique distinctive est bien visible : la dent cardinale la plus proche de la fossette ligamentaire est étroite et orientée vers le bas chez Bornia aartseni, alors qu’elle est plus épaisse et orientée vers l’arrière chez Bornia sebetia.

Autrefois considérée comme une sous-espèce de Bornia sebetia, cette coquille a été élevée au rang d’espèce en 2012 par S. Gofas.

4

Novapex / Société 14(1), 10 mars 2013

RÉCOLTES PERSONNELLES

Dans une laisse de mer récoltée en mai 20 1 2 sur la plage de Monte Gordo (Algarve - Portugal) (Carte - n°l), un petit spécimen double (Fig. 1) et 4 valves séparées de Bornia species (2 VG et 2 VD) ont été isolés d’un groupe de valves plus grandes appartenant à Bornia sebetia. Ces spécimens présentaient des caractéristiques en tout point identiques aux coquilles de Benalmâdena (Espagne - côte méditerranéenne) (Carte - n°3) : épaisseur et orientation d’une des dents cardinales de la valve gauche et taille des valves de la coquille.

Nous avons réexaminé tous nos lots de petits spécimens de Bornia récoltés en Algarve et en Andalousie atlantique.

Une valve gauche en provenance de Mazagon (Andalousie - Espagne - Mai 2003) (Carte - n°2) a également été identifiée

CONCLUSION

L’aire de distribution de Bornia aartseni est élargie de l’Andalousie méditerranéerme à l’Andalousie atlantique et aux côtes adjacentes de l’Algarve. La petite taille de l’espèce et sa proximité avec Bornia sebetia laissent à penser qu’elle est passée relativement inaperçue jusqu’à ce jour.

RÉFÉRENCES

Gofas, S. Moreno, D. & Salas, C. (coords.) 2011. Moluscos marinos de Andalucia. Mâlaga: Servicio de Publicaciones e Intercambio Cientifico, Universidad de Mâlaga. Volumen II, pp. i-xii y : 343-798.

Gofas, S. 2012. A New Species of Bornia (Bivalvia - Galeommatoidea) from Southern Spain. Iberus, 30(2): 41-48.

van Aartsen, J.J. 1996. Galeommatacea and Cyamiacea. La Conchiglia, 279: 31-36.

van Aartsen, J.J. 1996. Galeommatacea and Cyamiacea - Part IL La Conchiglia, 281: 27-53.

Novapex / Société 14(1), 10 mars 2013

5

L’écho des réunions

Marc ALEXANDRE

Réunion du 15 décembre 2012 (PC) Christiane Delongueville et Roland Scaillet : Richesse malacoiogique sur les côtes du détroit de Gibraltar

Christiane et Roland nous ont présenté une approche intégrée du détroit de Gibraltar. Plus qu’un lieu géographique mythique, ce détroit (14 Km dans sa partie la plus étroite) situé aux confins de l’Europe et de l’Afnque occupe une position privilégiée entre les deux continents. Le détroit est délimité en Atlantique par une ligne reliant le cap Trafalgar en Espagne au cap Spartel au Maroc et en Méditerranée par une ligne reliant la Punta de Europa à Gibraltar à la Punta Almina à Ceuta au Maroc Espagnol.

Pour en comprendre les particularités, il faut d’abord s’adresser à la géologie. Celle-ci nous apprend quand et comment s’est formé le détroit voici 5,6 millions d’années, lorsque la Méditerranée asséchée à l’époque et réduite à une cuvette d’évaporation s’est rouverte suite à la combinaison de différents facteurs qui ont amené à son remplissage en quelques dizaines d’années. La force colossale de l’eau pénétrant dans la cuvette méditerranéenne a sculpté le détroit tel que nous le connaissons à ce jour. Son histoire est écrite sur le profil de ses côtes et sur ses reliefs

sous-marins.

Les échanges d’eaux entre l’Atlantique et la Méditerranée ont lieu dans ce goulot d’étranglement. L’eau de l’Atlantique pénètre en surface, alors que par le fond, l’eau de la Méditerranée plus salée, plus chaude et plus lourde s’écoule vers les profondeurs de l’Atlantique. Il est donc nécessaire de nuancer la question de savoir où commence la Méditerranée et où finit l’Atlantique ou inversement. Les limites géopolitiques ou géographiques ne suffisent pas pour répondre à cette question. Les phénomènes hydrologiques qui se produisent dans le détroit sont donc à l’origine de la particularité de la faune malacoiogique que l’on y trouve.

Courants, températures de l’eau, salinité, richesse en nutriments expliquent comment se comporte cette faune située au point de rencontre des zones zoogéographiques lusitanienne, mauritanienne et méditerranéenne. Elle comprend des espèces endémiques (pour autant que des relevés futurs n’en agrandissent pas l’aire de répartition), des espèces méditerranéennes ne franchissant pas le détroit à l’ouest, des espèces atlantiques ne franchissant pas le détroit à l’est et enfin des espèces qui se retrouvent de part et d’autre du détroit, tant en Méditerranée qu’en Atlantique.

Christiane et Roland nous ont fait partager tous ces aspects du détroit à l’aide d’un magnifique Power Point qui a mélangé, cartes, graphiques, courbes bathymétriques, photos des lieux et photos des coquilles qu’ils y ont récoltées au cours de leurs nombreuses campagnes de collectes tant sur les côtes espagnoles que marocaines. Encore une fois, nos amis nous ont fait partager de merveilleux moments et qui sait peut-être donné l’envie d’aller voir par nous même la beauté et la richesse de ces magnifiques paysages.

Merci à vous deux.

6

Novapex / Société 14(1), 10 mars 2013

Morceaux choisis

Claude VILVENS & Etienne MEULEMAN

EZ3 Dissolutions calcaires

Cette coquille observée au rnicroscope électronique a balayage (à gauche) - appartenant au ptéropode Limacina helicina antarctica - a été prélevée à 200 mètres de profondeur dans l'océan Austral. Composée d'aragonite, forme de carbonate de calcium, elle montre déjà d'importants signes de corrosion comparée a une coquille intacte (a droite), comme vient de l'observer une équipe internationale. En cause ? L'acidification des océans, consécutive à la dissolution dans î'eair de mer dîme partie du dioxyde de carbone produit par les activités humaines. L'aragonite est en effet particulièrement sensible à l’acidité de i'eau de mer. A mesure que celle-ci augmente, la concentration des ions carbonate diminue, ce qui favorise la dissolution de l’aragonite, laquelle intervient dès 200 mètres. N Bedn3fsei<efc)/.,/V9eadoi:t0.1038/ngeol635,2012.

La Recherche 472

février 2013

Journal Métro du 8 janvier 2013

Athéna 287 janvier 2013

(Service public de Wallonie)

^ -k- ..

-fi.

TOKYO Des scientifiques et des chaînes de télévision japonaise et améiicaine ont annoncé hier avoir filmé pour la première fois un cala- mar géant par 900 m de fond dans l’océan Paci- fique. L’animtd inytlik|ue, de couleur argentée, a été filmé par une équipe du Musée scienti- fique national Japonais en collaboration avec: la chaîne de télévision publique japonaise NHK et la chaîne spécialisée Disœvery Channel. l£ calamar géant, dont le nom scientifique est ar- chiteutîîis, a été repéré par 630 mètres de fond par une équipe en sous-marin danslePadficiue nord. Le submersible a sum le géant jusqu’à 900 m de profondeur avant qu ’il ne disparaisse clans les abysses. lî? seul cor|3S du calamar a été évalué à 3 m de long. Sa longueur totale a été estimée à 8 m, en l’absence de ses deux princi- paux tentacuies qui étaient sectionnés . ■

Calamar géant dans le Pacifiqye

y e Glaacus a^lar-tlcm est un n -yllu^ttue gastéropode qui vit dans les eaux tem- t

nérées ou uopit.oî'-s dr monde '.ntier. Il floi^e juste en dessous de la surface de (

."e-v/, V7nî,-e ver\. '.•iur. H se noiunt prinâpaierrusnt d'hydrozoaires, par exemple ■ f /fl Physohks fyoir 4i o:na if 28^}, desquels il tire son pouvoir urticant. Ce bel animal marin es- Ueuo n- ';;c ^-'ue et mesme environ 35 mm.

Novapex / Société 14(1), 10 mars 2013

7

Quoi de neuf?

Claude VILVENS

Comme chaque année : la Bourse J'Antwerpen (Anvers) de la BVC !

BELGISCHE VERENIGING VOOR CONCHYLIOLOGIE [BVC] [ASSOCIATION ROYALE BELGE DE CONCHYLIOLOGIE]

www.bvc-gloriamaris.be

BOURSE INTERNATIONALE AUX COQUILLAGES

18 et 19 mai 2013 - Antwerpen - Belgium

Samedi 18 mai : 10 h - 18 h Dimanche 19 mai : 10 h - 16 h Admission : 2 Euros (-12 gratis)

Sports hall Extra Time

Louisalei 24

2660 Antwerpen - Hoboken

http://www.extratime.be

(tram 2, 4 en bus 1, 13, 33 tôt / descendre “Kioskplaats”)

5A ANTWERPEN CENTRUM

5 WiLRtJK

£>£ BRuyHtAAN

8

Novapex/ Société 14(1), 10 mars 2013

Quelques nouvelles publications

Roland SCAILLET

Maria Scaperroita Stoiaiio Bariolini Cesari; Bogi

ACCRESCIMENTl

STAni [)l ACCKhSCSMliKrO DFt MOU.L:saU MARlNi DEL MEDHEKKANED

-jt gîwîh wl' Uic IMM mtiîLwt'v oi îirt; Vlr(u!i2rr.:n«;jn Srw

Votunu- IV

Cela devient quasi une tradition. Chaque année voit paraître un volume supplémentaire d’un Opus commencé en 2009 ;

« Accrescimenti - Stadi di accrescimento dei molluschi marini del Méditerranée ».

2012 nous apporte donc le quatrième tome de la série. Ce nouveau millésime illustre 130 nouvelles espèces, ce qui porte à ce jour à 376, 131 et 1 respectivement le nombre de gastéropodes, bivalves et scaphopodes traités. Le principe reste toujours le même. En cinq ou six représentations, le portrait de l’espèce abordée est brossé, du stade juvénile à l’adulte. Cette démarche révèle parfois des surprises, tant les premiers stades de croissance d’un individu peuvent être différents de sa forme finale. Cela ravit à chaque fois ma curiosité de zoologiste et cela facilite également grandement mes activités de détermination comme collectionneur de petit matériel glané ça et là sur une plage, dans un port, ou sur le pont d’une barque.

Pour chaque taxon spécifique, la liste systématique des espèces illustrées à ce jour publiée en début d’ouvrage renvoie le lecteur au volume correspondant. Si l’Italien n’est pas votre langue maternelle, il n’y a aucun problème. La fiche technique se rapportant à chaque espèce est traduite en anglais en fin de volume.

Les auteurs l’annoncent, un cinquième tome est en préparation. Je me réjouis à l’avance de ce que la cuvée 2013 nous apportera. Maria Scaperrotta, Stefano Bartolini et Cesare Bogi, continuez donc à nous émerveiller, tant d’espèces restent encore à illustrer. Merci pour cette précieuse contribution à l’essor de la Malacologie.

Avis aux amateurs et bonne lecture.

Roland Scaillet

0La Société de Malflcologle • Mozilia flrefoK

Dctitef ËdWoQ A/6chage ti*stor<qij« Warau©-i>dgw Ç/utib î il ' c http //«www.societe-tKlge-de-nM^obgie.be/

vtivuiii R««.hw« IvîA >fad>jrMoriA . o<n e-5trvtcfi< 1 la Société Bnkje ffc M

Lâ Société 8ehie de Malacologie ^ A(f^9Sfori2010.DdI (Oblet .

U Société Balgif SS Vcliiala

La Société Belge de Malacologie > f

Accueil w

fSirmvtaw sur tr sitr ür la Sariélâ Hrlga ér Maiaralagir

Accueil

Mntluscjues

Clu/toy/aphieii

Réunions

Puhlicutions

Excursions

Hihliothiijue.

tn Sociétf Bïîge MaJar.'.îogjf' (eit la SBAft «t mr ïcicntiSfîUc en ASB! &âï)eof>h<'ric', cctK q»t soiü mtftrvsKS par :

• l'thrdc des moOmiptes (oiaiinf. teneshes et d'eau douce);

> Icia classifitalioti et Iciii ststémaiùpte, la colïecliou dev

l'etude et la coiuprclicnsioii des divers . dey iiioUitsqucs-

Expositions	a l'heure âchtcllc phw ou mÂ-ias 200xuén^rîî Actéa.awteunou prvfnsiofttt^ls Sci acfcx’Ues, bajces | vitlf! bènévoiert- «ont ■*55'5rrtJé9ern<ia reî r<îurii‘'>ns ifin a?énérai. «.te f ir avçr unr confér?r.fe sw un j
Conseil	rnéthir ses wk .r 'fzin n (Viiwmrnmi 2 à .3 pir .nq. ra ■ ptibjw'aijw3î réguitm dm rnitnéros bor,*; jjfhr. |
	Jr;! siui’Uï iiiî Ir-t rij^Bu^uu» icrrt'iirci vt diiL'. tic:’ pul-liLaü^-<rj dr/ascs) airiix q«\mtr îri<î.>«>sia<'u tasiueStr cl WM/bouist |
Membres	•9C''3a<‘nfteI«î j
Annonces	Piochauîc avlivitc . 1
X Raclaîiüie' ;	f J E.e*4H.'t.tCT la
Terrra^

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Novapex / Société 14(1), 10 mars 2013

9

Nous avons reçu

Etienne MEULEMAN

4 Le coin du Débutant par G. Jaux

7 Voyage aux Iles Marquises par D. Wimart-Rousseau et P. Marti

10 Retour de vacances en Corse par D. Marcellesi

11 The protoconch and early teleoconch whorls of the brownish cerithiopsis in the Caribbean and adjacent région by E. Rolân, H. G. Lee, M. F. Krisberg & R. Femândez-Garcés

17 Le genre Neritopsis, un fossile vivant par F. Vigne

18 Cypraea stercoraria star des Cypraeidae de l’Ouest Afrique par J-F. Michard

21 Coup de projecteur sur les Nérites par L. Limpalaër 26 Les trois cônes de Méditerranée par J-P. Sidois 28 Les Volutes fossiles de France par D. Ledon 30 Les ancêtres de Xenophora (2®"’^ partie) par N. Laurenceau

35 Bourse de Rennes 2012 - Petites annonces

36 Nains et Géants

39 Echo...quillages

XENOPHORA

(France)

N°139, juillet-août-septembre 2012

SPIRULA

(Pays-Bas)

N° 387, juillet-août 2012

Diverse bronnen Excursies en Malacologische agenda Ncdcrland 93

Redactie Erratum - Spirula 385 pagina 48 94

E. Gittenberger De moléculaire revolutie in de systematiek: zin en onzin 95

A. W. Janssen Pteropoden en oceaanverzuring 96

D. Leeuw Zeebiologie, zeeaquariologie en malacologie

Een geschiedenis van Biologia Maritima die Icidde tôt de Spirula

zoals we die nu kennen 98

EJ. van Gcmert Pcnang en de Arcidac van Jousscaume - Penang and the

Arcidae of Jousseaume 105

Nicuw in Nederland (samcngcstcld door G.D. Majoor) 107

B. Goetheer Fleeft de Filippijnse tapijtschelp Ruditapes philippinamm

(Adam & Rccvc, 1850) zich in de Westerschelde gevestigd? 107

W. Faber Nieuw beschreven mariene molluskensoorten - (new taxa:

marine molluscs) 108

W. Faber Artikelen in tijdschriften - (journal papers; marine malacology) 111

W. Faber en J. Goud Nieuwe boeken - new books 117

W. Faber Schelpcnbeurzcn en bijeenkomsten 120

10

Novapex / Société 14(1), 10 mars 2013

CONCHYLIA

(Allemagne)

N°42, 1-4, mai 2012

Conchylia

Inhalt von / Contents of 42 (1-4), Mai/May 2012

Inhalt/Contents 1

Kronenberg, g. C.: Identifications for Stromboidea (Caenogastropoda) in the facsimile reprint of the De Favanne de Montcervelle 1780 édition of “La Conchyliologie, ou Histoire naturelle . . by DÉzallier d’ Argenville 3

Egorov, R. V.: Conchological Review of the Land Snails of the Genus Caucasigena Lindholm, 1927 (Gastropoda: Pulmonata: Hygromiidae) with Some Notes on Other Caucasian Hygromiids 13

Salisbury R. A., Herrmann, M. & Dekkers, a. M: Description of a new

red-spotted Costellarid (Gastropoda: Costellariidae) from the Indo-Pacific with remarks on Vexilliim (Costellaria) unifasciatum (WOOD, 1828) and Vexillum {Costellarid) clathratum (Reeve, 1844) 27

Herrmann, M. & Gori, S.: Two new species of Mitromica and Thala (Gastropoda:

Costellariidae) from Oman and Mozambique 39

Fehse, D.: Caribeginella ESPINOSA & Ortea 1998 eine Gattung der Tivioidea

(Mollusca: Gastropoda: Littorinimorpha)? 49

Morrison, h. m.: Descriptions of two new species of Amoria (Gastropoda:

Volutidae) from Western Australia 51

Lorenz, F.: Another New Species of Archivolva from the Red Sea (Gastropoda: Ovulidae) 65

Wiese, V., Richling, I., Brinkmann, R. & Groh, K.: Weichtier des Jahres 201 1 :

Die Zierliche Tellerschnecke Anisus vorticulus 7 1

Nordsœck, h.: Die Medora-AsX^n Italiens (Gastropoda, Stylommatophora, Clausiliidae, Alopiinae), mit Beschreibung einer neuen Unterart von Medora dalmatina Rossmàssler 75

Lorenz, F.: Nesiocypraea midwayensis kontiki n. ssp., a New Subspecies from the Eastem

Pacific (Gastropoda: Cypraeidae) 83

Lorenz, F.: A New Name for Are storide s argus contrastriata (Perry, 1811)

(Gastropoda: Cypraeidae) 87

Huber, m.: Donax bruneirufi and Donax baliregteri, Two New Deltachion from the

Central Indo-Pacific (Bivalvia: Donacidae) 97

Nordsieck, R.: Erstaunliches von den Schnegeln (Limacidae Rafinesque, 1815) 105

Kittel, K.: Description of Theba lindneri n. sp. from Fuerteventura, Canary

Islands, Spain (Gastropoda, Helicidae) 1 1 1

Fehse, D.: Description of a New Barycypraea SCHILDER, 1925 from the Indonesian

Miocene (Mollusca: Cypraeidae) 117

Novapex / Société 14(1), 10 mars 2013

NOTICIARIQ DE LA SQCIEDAD

ESPAnQLA de MALACQLQGIA

(Espagne)

N°57, 2012

INDICE

Editorial

Secretaria ;

Tesorerîa

Informe del Editor

Recensiones Bibliogrâficas

Noticias Malacolôgicas

Actualidad legislativa y malacologia

Taller de fotografia de moluscos

Queremos saber

Colaboraciones

- Sansonia tuberculata (Gastropoda, Pickworthiidae) en Cuba: atgunos nuevos datos

■ Hastula Aciculina (Gastropoda, Terebridae): Nuevas Aportaciones

- Sobre el hallazgo de una nueva poblacion en Espana de Quickella arenaria

(Potiez & Michaud, 1835) (Gastropoda: Succineidae)

- Moluscos terrestres de las islas e islotes del litoral de la Région de Murcia y Mar Menor

(Sureste de la peninsula Ibérica)

- Melanopsis Tricarinata (Bruguière, 1789) (Gastropoda. Melanopsidae) en Navarra (Espana)

- Primera Cita de Triîonia Lineata Aider y Hancock, 1848 (Gastropoda; Tritoniidae)

en Andaluct'a, Sur de la Peninsula Ibérica

- Primera Cita de Tritoniopsis Cincta (Pruvot-Fol, 1937) (Gastropoda: Opisthobranchia;

Tritoniidae) en Andalucia, Sur de la Peninsula Ibérica

Preguntas a... Luis Murillo •

Indices de Revistas

Pasatiempos

3

4

5

10

11

16

36

50

51

52 55

57

62

67

75

77

79

82

98

IBERUS

(Espagne)

Vol. 30, N° 2, juillet 2012

Bogi C., Karhan S.Ü. and Yokes M.B. Oscilla galilae, a new species of Pyramidellidae (Mollusca, Gastropoda, Heterobranchia) from the Eastem Mediterranean 1-6

Holoyoak D.T., Holoyoak G.A. and Torres Alba J. S. A reassessment of the species of Truncatellina (Gastropoda : Vertiginidae) in the Iberian Peninsula and North-west Africa 7-33

Rolân E. and Rubio F. A new species and range extension of Ponderinella (Gastropoda, Tomidae) in West Africa 35-39

Gofas S. A new species of Bornia (Bivalvia : Galeommatoidea) from Southern Spain....41-48

Oliver J.D., Templado J. Y Kersting D. Marine gastropods of the Columbretes Islands (Western Mediterranean) 49-87

Templado J. and Rolân A new species of Phorcus (Vetigastropoda, Trochidae) from the Cape Verde 89-96

Lefrere L., Moukrim A., Idardare Z., Bergayou H. and Kaaya A. Reproductive cycle of Scrobicularia plana (da Costa, 1778) (Bivalvia : Semelidae) in two Moroccan lagoons ; Khnifiss and Oualidia 97-106

Antit M. et Azzouna A. Mollusques des milieux littoraux de la baie de Tunis 107-133

12

Novapex / Société 14(1), 10 mars 2013

MQLLUSCAN RESEARCH

(Australie)

Vol. 32,N°2, mai 2012

Molluscan

Research

An international jonnial of l'Iie Malaeolooical Society of Aiislralasia The Society for tlie Study of Molluscan Diversity \ olunie 32, .Nuniher 2 (May 2012)

Tonnoidean gastropods of French Polynesia

A.G BEU, P. BOUCHET & J. TRÔNDLÉ

VISAYA

(Philippines)

Vol. 3, N° 6, octobre 2012

03 ABOUT VISAYA

04 Taxonomie Review of the Conus spectrum.

Conus stramineus and Conus coUisus Com- plexes (Gastropoda: Coinidae) Part 111: The Conus collisus Complex

R. M. (MIKE) EILMER

49 Pionoconus rohini (Gastropoda, Conidaf.) New' Spccies from the South Western Philippines.

E. MMPALAËR & E. MONNIER

55 One New Species of Capulus from l'aiwan with Remarks on C. japonicus A. Adams, 1861 and C. violaceus (Angas, 1867) (Gastropoda: Capiji.idae)

SHIli-l tlUANG& YA-FAN HUANG

61 Notes on the Taiwane.se Fossarinae Species with the Description of One New Species (Gastropoda: Pi.aximdae)

SHIH-I HUANG

65 A .Statement on the Uand Snails of Cebu

GUlOO T. POPPE, SHEILA P. TAGARO, DANIEL CROOKSHANKS, KLAUS GROH & JERLYN SARINO

Novapex / Société 14(1), 10 mars 2013

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Pree list on reguesr, gotxi quaiity shcîls at dic bcst pnce/t. Satisfaction /

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Novapex / Société 14(1), 10 mars 2013

Dutch

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Our society warmiy welcomes new members {both from the Netherlands and abroad) to participate in our activities:

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Join us and ineel new shelling friends. Further info; Bram Breure, Van Schagenplanlsoen 8, NL-2741 EN Waddinxveen, The Netherlands. E-mail: abreure(3>x.s4all,nl

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16

Novapex / Société 14(1), 10 mars 20 1 3

Grandes marées de l’année 2013

Christiane DELONGUEVILLE et Roland SCAILLET

2013 est une année plus que médiocre en matière de grandes marées. La plus grande n’a qu’un coefficient de 109. Par contre, il y a beaucoup de grandes marées modestes tout au long de l’année et ce y compris en janvier et en décembre.

Coefficients (> 100) des pleines mers à Brest

(Les marées basses correspondantes sont donc particulièrement intéressantes à prospecter)

Janvier	Samedi 12	102 - 104
	Dimanche 13	106- 106
	Lundi 14	104-102

Juin	Lundi 24	102-104
	Mardi 25	105 - 104
	Mercredi 26	103 - 100

Février	Dimanche 10	100-104
	Lundi 1 1	106-107
	Mardi 12	106-104
	Mercredi 13	101 - (97)
	Jeudi 28	100 -(99)

Mars	Lundi 1 1	(97) - 100
	Mardi 12	102-103
	Mercredi 13	103 - 101
	Mercredi 27	(96)- 100
	Jeudi 28	103 - 104
	Vendredi 29	105 - 104
	Samedi 30	102 -(99)

Juillet	Mardi 23	102-105
	Mercredi 24	107 - 108
	Jeudi 25	107-104
	Vendredi 26	100 -(95)

Août	Mercredi 2 1	103 - 106
	Jeudi 22	108-109
	Vendredi 23	108-105
	Samedi 24	101 -(96)

Septembre	Jeudi 19	101 - 104
	Vendredi 20	105 - 105
	Samedi 21	104-101

Octobre

Novembre Lundi 4 (99) -101

Mardi 5 101 - 100

Beaucoup de marées intéressantes auront lieu durant le week-end. Il est à espérer que les pêcheurs à pied seront plus respectueux de l’environnement que ce que nous avons pu observer l’année passée en Bretagne.

Conseils pour une marée réussie et soucieuse de l’environnement : Remettez toujours les pierres déplacées en bon ordre. Observez, photographiez et n’échantillonnez que le strict nécessaire. Soyez prudents et renseignez-vous sur les heures des marées à l’endroit où vous vous trouvez.

REFERENCE :

Annuaire des Marées pour l’année 2013 - Ports de France - Métropole - Tome 1 - SHOM (Service Hydrographique et Océanographique de la Marine) - Paris - 257 p.

Pointe de Kerpenhir - Locmariaquer (Morbihan)

Les données reprises dans cet article peuvent également se retrouver sur notre site Internet :

http://www.societe-belgc-de-malacologie.be

Décembre	Mardi 3	(98)- 101
	Mercredi 4	102-103
	Jeudi 5	102- 100

Avril	Vendredi 26	103 - 105
	Samedi 27	106-105
	Dimanche 28	104-101

Mai	Samedi 25	(99)- 102
	Dimanche 26	104-104
	Lundi 27	104- 102

N#VAPEX

MCZ

LIBRARV

JUL 1 0 2013

HARVARD

Trimestriel de la Société Belge de Malacologie UNI VERSIT'

association sans but lucratif

Quarteriy of the Belgian Malacological Society

VOL. 1 4 (2)	2013	10 JUIN
	SOMMAIRE
	Articles originaux - Original articles
R. Houart	Revised classification of a group of small species of Cytharomorula Kuroda, 1953 (Muricidae: Ergalataxinae) from the Indo-West Pacific	25
R. Goethals & D. Monsecour	A new species of Rolaniconus (Gastropoda: Conidae) from the Philippines	35
K. Fraussen & P. Stahlschmidt	Eclectofiisus, a new genus for Pararetifusus dedonderi Fraussen & Hadom, 2006 (Gastropoda: Buccinidae)	39
R. Houart & M. Severns Description of a new species of Favartia {Pygmaepteiys) (Gastropoda: Muricidae: Muricopsinae) from Hawaii	43
	NOVAPEX/SOCIETE
C. Vilvens	Prochaines activités (et annonce du banquet de la SBM)	17
C. Delongueville & R. Scaillet	- ^ Mollusques épibiontes et parasites sur venosa (Valenciennes, 1846) en Mer de Mamiara	19
C. Delongueville & R. Scaillet	Espagne, plage de Tarifa le 26 février 2013 : ^ laisse de mer réduit à 2 g de coquilles à détenniner	24
(suite du sommaire en dernière page de couverture)

ISSN 1375-7474

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- APEX (1986-1999)

SOCIETE BELGE DE MALACOLOGIE

R. Houart

Novaphx 14(2): 25-34, 10 juin 2013

Revised classification of a group of small species of Cytharomomla Kuroda, 1953 (Muricidae: Ergalataxinae)

from the Indo-West Pacific

Roland HOUART

MCZ

LIBRARN

JUL 1 0 201

HARVARI

UNIVERS!"

Research associate

Institut royal des Sciences naturelles de Belgique, rue Vautier, 29, B- 1000 Bruxelles, Belgium

roland. houart (^skynet.be

Keywords. Gastropoda, Muricidae, Ergalataxinae, Cytharonionila, Indo-West Pacific.

Abstract. Five similar looking species of Muricidae from the Indo-West Pacifie are reviewed, illustrated and commented: Cytharomomla amhonensis (Houart, 1996), C. henedicta (Melvill & Standen, 1895), C. dollfusi (Lamy, 1938), C. lefevreiana (Tapparone Canefri, 1880) ) and C. paiicimacidata (Sowerby, 1903). The type material is illustrated for ail the species. A lectotype is designated for Cytharomomla lefevreiana. The radula morphology is described.

Résumé. Cinq espèces de Muricidae morphologiquement très proches, vivant dans l'Indo- Pacifique oceidental, sont révisées, illustrées et commentées: Cytharomomla amhonensis (Houart, 1996), C. henedicta (Melvill & Standen, 1895), C. dollfusi (Lamy, 1938), C. lefevreiana (Tapparone Canefri, 1880) et C. paucimacidata (Sowerby, 1903). Le matériel type est illustré pour toutes les espèces. Un lectotype est désigné pour Cytharomomla lefevreiana. La moiphologie de la radula est décrite.

INTRODUCTION

A small group of tiny muricid species occumng in the Indo-West Pacific and here reviewed ineludes only five species, but these were onee classified in several généra, occasionally misidentified, and are still little known, except for a more eommon speeies found mainly in the Philippine Islands but occumng worldwide.

None of these species is really rare except Cytharomomla paucimacidata from Japan of which few specimens are cuirently known. In this group of speeies not only the genus, but also the synonymy differs between published works. That was the reason for studying this group very carefully and to elear up its classification.

Ail the types were examined as well as an impressive number of specimens studied throughout the years.

The terminology used to deseribe the spiral cord configuration follows Merle (1999, 2001) and is illustrated in Figs 4-6. PI: shoulder cord; P2-P6: primary cords of the convex part of the teleoeonch whorl.

Material and mcthods

Much of the material examined was eolleeted during the expéditions eonducted by MNHN and IRD (formerly ORSTOM) in the Tropical Indo-Pacific. Specimens were eolleeted during following eruises: MD32 (1982) in Reunion, MONTROUZIER ( 1993) in New Caledonia, MUSORSTOM 9 (1997) in The Marquesas Islands, SUVA 4 (1999) in Fiji, ATELIER

LIFOU (2000) in New Caledonia, PANGLAO 2004 (2004) in the Philippines, SANTO (2006) in Vanuatu, SALOMONBOA 3 (2007) in the Solomon Islands and ATIMO VATAE (2010) in Madagascar. Other studied speeies belong to the author’s eollection. Ail the specimens are listed and were examined by the author. No mention of living or empty shell is given, mostly because in many cases it was quite impossible to décidé whether the shell was eolleeted dead or alive, the living specimens having their operculum deeply retracted inside of the shell. However, the species live in depths of 3-20 m, except C. lefevreiana for which specimens, rarely eolleeted alive in the examined samples, were dredged in 55-73 m and C. dollfusi which is usually dredged in tangle nets with shell grit, in 50-150 m in the Philippine Islands.

The eomplete data is given for many stations of the MNHN-IRD expéditions, but when there is an extensive list of stations only the name of the expédition and the number of the station is given. The number of examined specimens is mentioned between brackets for all the samples.

Additional references are added only when it concems a misidentification.

Abbreviations

IRD: Institut de Recherche pour le Développement. MNHN: Muséum national d'Histoire naturelle, Paris, France.

MZUF: Museo di Storia Naturale, Sezione di Zoologia "La Speeola", Université di Firenze, Italy.

NHMUK: Natural History Muséum, London, United Kingdom.

25

R. H ou A RT

Classification of small species of Cythcironwnila.

ORSTOM: Office de la Recherche Scientifique et Technique Outre-Mer.

RH; collection of the author.

RMN H; Nationaal Natuurhistorisch Muséum, Leiden, the Netherlands (now in Naturalis Biodiversity Center, Leiden, the Netherlands).

Discussion

In a recent paper (Claremont et al, 2013), a subclade contains nominal species of Pascula, Orania and Cytharomonila. Based on the species included in their study, ail these généra, as currently defmed, are polyphyletic. It is possible that the entire clade should be assigned to one genus, but beeause the oldest name is Orania (type species not yet included in the analyses), followed by Pascula (type species not yet included in the analyses), then Cytharomonila, the authors were unable to detennine which name to propose. Pending a complété révision of this subclade, they conservatively retained ail species in the généra to which they were previously assigned. Only C. doUfusi is included in their molecular analyses (as Cytharomonila paucimaciilata). This

misidentification is strictly my fault beeause this was the name I used for the Philippine shell before the présent review was undertaken.

My decision to maintain these species in Cytharomonila while one or more species included in the présent study were once classified in Pascula (Cemohorsky, 1982, Trôndlé & Houart, 1992, Houart, 1995, Higo, Callomon & Goto, 1999, Tsuchiya, 2000, Mienis, 2003, Heiman & Mienis, 2003, Houart, 2008) or in Cytharomonila (Houart, 2008, Houart & Trôndlé, 2008, Houart & Héros, 2008, Houart & Gofas, 2012, Claremont et ai, in press), is also based on this molecular study. I decided to retain these species in Cytharomonila as was done in the more recent papers.

The most recent classification known of these species was submitted as follows by Houart and Gofas (2012) (WoRMS):

Cytharomonila amhonensis (Houart, 1996). Cytharomonila lefevreiana (Tapparone Canetri, 1880)

=Murex (Ocinebra) benedictus Melvill & Standen, 1895.

Cytharomonila paucimaciilata (Sowerby, 1903)

= Murex dollfusi Lamy, 1938.

The Lipdated classification after examination of the type material and of specimens originating from throughout their geographical distribution looks now as follows:

Cytharomonila amhonensis (Houart, 1996). Cytharomorula henedicta (Melvill & Standen, 1895). Cytharom or nia dollfusi (Lamy, 1938). Cytharomorula lefevreiana (Tapparone Canefri, 1880).

Cytharomorula paucimaciilata (Sowerby, 1903).

SYSTEMATICS

Family MURICIDAE Rafmesque, 1815 Subfamily ERGALATAXINAE Kuroda, Habe & Oyama, 1971

Genus Cytharomorula Kuroda, 1953

Type species by monotypy: Cytharomorula vexillum

Kuroda, 1953, Japan.

Description of this group. Shell small, up to 12 mm in length at maturity, lanceolate, narrow, nodose. Subsutural ramp broad, strongly sloping. Spire high with 3. 5-4. 5 protoconch whorls and teleoconch of up to 5 weakly convex, shouldered, nodose whorls. Protoconch conical, with sinusigeral notch, whorls smooth (Fig. 7).

Axial sculpture of last teleoconch whorl consisting of 7-10 low or moderately high ribs and varices, crossed by primary, secondary and tertiary spiral cords, P1-P3 high, broad and obvions, P3 usually strongest, P4 smaller, narrow, P5 and P6 very small or obsolète. Intersection of axial and spiral cords giving rise to small nodes or broad, low, open spinelets.

Aperture small, ovate. Columellar lip narrow, occasionally with 2 or 3 knobs abapically, with low pariétal tooth at adapical extremity. Outer lip with weak or strong, occasionally split, broad or elongate denticles within. Siphonal canal very short, dorsally recurved, with P5 and P6.

Operculum light brown, ovate-elongate with subapical nucléus.

Three-dimensional radula (Figs 1-3) with rachidian tooth bearing a long, narrow, prominent central cusp, a single small latéral denticle, a short, relatively broad latéral cusp and occasionally few marginal folds. Latéral tooth sickle-shaped with broad base.

Cytharomorula amhonensis (Houart, 1 996)

Figs 2-3,4, 8-9, 14-19

Pascula ambonensis Houart, 1996: 383, figs 17-18.

Additionai référencé

Pascula lefevreiana - Houart, 2008: 204, pl. 397, figs la, Ib (not Tritonidea lefevreiana Tapparone Canefri, 1880).

Type material. Holotype RMNH 57155.

Type locality. Indonesia, Ambon Id, Hitu, Suli, in sand, near coral.

Other material examined. Mozambique:

Quissimajula, N Mozambique, 2-3 m, in sand and coral, RH (4); Nacala Bay, dived in 2-4 m, RH (6 ); Nacala Bay, dived in 2-3 m, RH (3); Feifiao Veloso Bay, 3-4 m, in dead coral, RH ( 1 ); Madagascar. Tulear, MNHN (2); ATIMO VATAE, stn TBOl, MNHN (1); stn TS02, MNHN (1); stn TS03, MNHN (1); stn TS04, MNHN

26

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Novapex 14(2): 25-34, 10 juin 2013

(1); stn TV07, MNHN (2); sln TS17, MNHN (1); stn BP4I, MNHN (1); Maldives: Maagau Kandu, Ari Atoll, 20-25 m, RH (1); Papua New Guinea: Hansa Bay (Madang Province) Laing Island, RH (1); Philippines: PANGLAO 2004, stn SI, MNHN (4); stn B5, MNHN ( 1 ); stn B7, MNHN ( 1 Iv); stn B 1 0, MNHN (1); stn B 13, MNHN; stn S32, MNHN (1); stn B37, MNHN ( 1 ); stn B39, MNHN (2 ); stn L50, MNHN ( 1 ) Vanuatu: SANTOS, stn ZB06, MNHN (3); stn DB08, MNHN (3); stn ZB09, MNHN (4); stn FS51, MNHN

(1) ; stn FB92, MNHN (1); New Caledonia: ATELIER LIFOU, Loyalty Islands, Lifou, stn 1457, MNHN (8); 20°47' S, 167°03' E, 5-10 m, MNHN IM-20 10-21 107

(2) ; Marquesas: Motu One, MUSORSTOM 9, stn DW 1 29 1 , 7°48' S, 1 40°2 1 ’ W, 450-455 m, MNHN ( 1 ).

Distribution. Mozambique, Madagasear, Maldives, Indonesia (Ambon Island and Papua New Guinea), Philippines, Vanuatu, New Caledonia and French Polynesia (Marquesas).

Remarks. Cytharomomla ambonensis is a small, spiny speeies reaehing 6-10 mm in length. It differs from C. benedicta in having a more spiny appearanee and in having a relatively larger shell for the same number of teleoconch whorls, reaehing a length of 10 mm VS 8 mm in C benedicta', the primary spiral cords are broader and higher, moreover, P1-P3 are of an almost similar strength in C. ambonensis while PI and P2 are much naiTower and lower than P3 in C. benedicta', the secondary spiral cords between PI and P2, and P2 and P3 are broader and less numerous, 2 or 3 cords in C. ambonensis vs 3-6 in C. benedicta', the primary spiral cords P5 and P6 are also broader and higher in C. ambonensis while the P4 cord is smaller but still much stronger than the P4-P6 cords in C. benedicta', the area between P3 and P6 in C. ambonensis is also less concave than in C. benedicta. The colour also differs, being whitish with some dark brown secondary spiral cords and occasionally dark blotches on the primary eords in C. ambonensis vs having brown blotches on the subsutural ramp and between P3 and P6 in C. benedicta.

Cytharomomla ambonensis is sympatric with C. benedicta and C. paucimacidata in New Caledonia and Guam and with C. paucimacidata in the Philippines.

Cytharomorula benedicta (Melvill & Standen, 1895) Figs 1, 6-7, 20-26

Murex {Ocinebra) benedictus Melvill & Standen, 1895: 108,pl. 2, fig. 13.

Additioual refereuces

Pascula lefevreiana — Houart, 1995: 274, fig. 142; Tsuchiya, K. 2000: 385, fig. 105 (not Tritonidea lefevreiana Tapparone Canefri, 1880).

Cytharomomla lefevreiana — Flouart, 2011: 280, fig. 6 (not Tritonidea lefevreiana Tapparone Canefri, 1 880).

NOT Pascula benedicta — Cernohorsky, 1982: fig. 20 only (= Tritonidea lefevreiana Tapparone Canefri, 1880).

Type material. Lectotype Manchester Mus. EE.3708, (originally figured specimen. See also remarks) and 18 paralectotypes, Manchester Mus. E.7667, EE.7668, EE.7669and EE.8115.

Type locality. Lifou, in shell-sand.

Other material examiued. New Caledouia.

EXPEDITION MONTROUZIER, Touho, stn 1237, MNHN (1); stn 1240, MNHN (2); stn 1245, MNHN (7); stn 1255, MNHN (2); stn 1259, MNHN (7); stn 1260, MNHN (1); stn 1261, MNHN (4); stn 1268, MNHN (1); stn 1269, MNHN (23); stn 1270, MNHN (7); stn 1271, MNHN (12); stn 1273, MNHN (7); Koumac, stn 1302, MNHN (2); stn 1308, MNHN (1); stn 1310, MNHN (1); stn 1311, MNHN (5); stn 1312, MNHN (5); stn 1315, MNHN (5); stn 1316, MNHN (26); stn 1318, MNHN (18); stn 1319, MNHN (17); stn 1331, MNHN (16); stn 1333, MNHN (1); ATELIER LIFOU, Loyalty Islands, Lifou, stn 1413, MNHN (1); stn 1420, MNHN (9); stn 1421, MNHN (2); stn 1422, MNHN (1); stn 1423, MNHN (3); stn 1425, MNHN

(7) ; stn 1427, MNHN (4); stn 1429, MNHN (16); stn 1430, MNHN (1); stn 1432, MNHN (26); stn 1434, MNHN (2); stn 1432, MNHN (15); stn 1436, MNHN

(8) ; stn 1438, MNHN (1); stn 1441, MNHN (4); stn 1442, MNHN (4); stn 1443, MNHN (2); stn 1444, MNHN (4); stn 1446, MNHN (3); stn 1449, MNHN

(3) ; stn 1450, MNHN (7); stn 1451, MNHN (21); stn 1453, MNHN (4); stn 1454, MNHN (14); stn 1455, MNHN (1); stn 1456, MNHN (3); stn 1457, MNHN

(4) ; stn 1459, MNHN (2); Lifou (without any other data), MNITN (58); Touho, 15-20 m, RH (5); Koumae, 15-20 m, RH (2); Vauuatu. SANTO, stn DS06, MNHN

(1) ; stn ZB06, MNHN (1); stn DB08, MNHN (1); stn ZB09, MNHN (1); stn DB12, MNHN (2); stn ZB20, MNHN (1); stn DB33, MNHN (1); stn DB63, MNHN

(2) ; sta FB64, MNHN (2); Guam. W of Cocos Island, among dead coral, 19-28 m, RH (1).

Distribution. New Caledonia (Lifou and Koumac), Vanuatu, Guam, Japan (Kii Peninsula and southward) (Tsuchiya, K. 2000, as Pascula lefevreiana), French Polynesia (Tuamotu Arehipelago) (Cernohorsky, 1982), and Hawaii (Houart, 2011, as Pascula lefevreiana).

Remarks. This speeies is only kiiown from a few localities but it probably lives in other parts of the Western and Central Pacifie, maybe overlooked due to its small size and less known siibfamily. It was confused with and illustrated as C. lefevreiana in a few publication. See under C. lefevreiana for a comparison with that speeies.

Cernohorsky (1982: 128), in a study of some Indo- Pacifie Mollusca, classified C. benedicta in Pascula, he illustrated the leetotype of C. benedicta (Cernohorsky,

27

R. HOUART

Classification of small species of Cytharomonila.

1982: figs 12-13) and the holotype of C. paucimaculata (figs 18-19) but he synonymised both names. He figured the lectotype of C. beuedicta as being the holotype, but no holotype was designated by the authors, while there are 1 8 other "possible" syntypes in the Manchester Muséum (Machin, in litt.). In fact, Melvill & Standen (1895; 108) noted "A very small but exquisite shell of which we hâve only seen one full- grown specimen". This sentence implies that there were other specimens examined by the authors but that none of those were ftill-grown.

Therefore, in aecordance with ICZN (Art. 74.6) the specimen figured by Cemohorsky (1982) as being the holotype is deemed to hâve been designated as the lectotype (Figs 20-21).

Cemohorsky (1982: fig. 20) also illustrated a slenderer

specimen from an unknown locality, from the National Muséum of Wales. This specimen is not C. henedicta but C. lefevreiana which is narrower and slenderer with lower apertural denticles, fewer cords on the subsutural ramp, a lower and less obvions P3 spiral cord and a less concave area between P3 and P6 as observed in Cemohorsky's figure 20. Cemorhorsky's figure 21 on the same plate is another species currently known as Pascula darrosensis (E.A. Smith, 1884).

The shell illustrated by Trôndlé & Houart (1992: 87, fig. 45) is misidentified, see Houart & Trôndlé (2008: 71). Nevertheless, the presence of C. benedicta in French Polynesia is confimied by Cemohorsky (1982: 128, figs 14-15).

Also see under C. paucimaculata for a comparison with that species.

Figures 1-3. Radulae

1. Cytharomonda benedicta (scale bar 50 pm); 2-3. Cytharomonila ambonensis (Houart, 1996) (scale bars 100 pm).

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NOVAPEX 14(2); 25-34, 10 juin 2013

Cytharomonila dollfusi (Lamy, 1938)

Figs 33-40

Murex dollfusi Lamy, 1938: 54, fig. 1.

Additional reference

Cytharomonila paucimaculata — Houart, 2008: 198, pl. 394, figs 2-5; Houart & Héros, 2008; 459 (not Peutadacty’lus paucimaculatus Sowerby, 1903).

Type material. Holotype MNHN 0103.

Type loeality. Jubal Island (Juzur Jabal, 27°40' N, 33°48’ E, Egypt).

Other material examined. Madagascar. ATIMO VATAE, stn DW3606, MNHN (1); Maldives. Maagau Kandu, Ari Atoll, 20-25 m, RH (1); Philippines. PANGLAO 2004, stn Pl, MNHN (3); stn SI, MNHN (12); stn R3, MNHN (3); stn P4, MNHN (64); stn B7, MNHN (2); stn SI, MNHN (11); stn Tl 1, MNHN (3); stn BIO, MNHN (7); stn B12, MNHN (2); stn B14, MNHN (1); stn B 15, MNHN (2); stn B 16, MNHN (4); stn B 17, MNHN (2); stn B 19, MNHN (2); stn B21, MNHN (2); stn B24, MNHN (2); stn B25, MNHN (1); stn B28, MNHN (1); stn S28, MNHN (6); stn SI, MNHN (1); stn B32, MNHN (2); stn B36, MNHN (5); stn B37, MNHN (4); stn B38, MNHN (2); stn B39, MNHN (7); stn B40, MNHN (4); stn B41, MNHN (3); stn L41, MNHN (1); stn B42, MNHN (10); stn L42, MNHN (2); stn L46, MNHN (29); stn L50, MNHN (3); stn L51-60, MNHN (27); stn L65-68, MNHN (5); stn L69-73, MNHN (14); stn L74-75, MNHN (9); stn L76, MNHN (213); stn L79, MNHN (11). Between Bohol and Cebu, 50-100, local fishermen, MNHN (117); Panglao, 80 m, RH (13); Cebu, Mactan Island, Punta Engano, in tangle nets, RH (43); Balut, RH (1); Between Bohol and Cebu, 50-150 m, RH (3); Bohol, tangle nets, RH (9); Sulu Sea, RH (61); Balicasag Island, 160 m, rocky sand bottom, RH (1); S Mindanao, Aliguay, Dipolog, 150-180 m, tangle nets, RH (8); New Caledonia. ATELIER LIFOU, Loyalty Islands, Lifou, stn 1435, MNHN (1); stn 1465, MNHN (1); Vanuatu. SANTO, stn ZB13, MNHN (1); stn FB68, MNHN (1); stn DS99, MNHN (1); stn DS103, MNHN (9); mixed, MNHN (1); Solomon Islands. W San Cristobal, SAEOMONBOA 3, 10°25’ S, 161°22' E, 121-180 m, MNHN (1); Fiji. SUVA 4, stn DW 09, 18°2rS, 178°06’E, 37-41 m MNHN (1); stn DW 12, 18°2rS, 178°10’E, 39 m, MNHN (1); stn DW 25, 18°27’S, 178°01’E, 48-51 m, MNHN (1); French Polynesia. Marquesas, MUSORSTOM 9, stn DW 1148, MNHN (1); stn DWl 154, MNHN (3); stn DW1170, MNHN (l);stn CP 1 177, MNHN (1); stn CPI 178, MNHN (1); stn DR 11 82, MNHN (2); stn DR 1200, MNHN (8); stn

DW 1204, MNHN (8); stn DW 1208, MNHN (1); stn

DW 12 10, MNHN (3); stn DR 1223, MNHN (1); stn

DW 1224, MNHN (3); stn CP 1228, MNHN (1); stn

DW 1242, MNHN (2); stn DR1315, MNHN (3);

Austral Archipelago, BENTHAUS, stn DW 1869, 28°58' S, 140° 15' W, 40-440 m, MNHN (2); DW 1926, 24°38' S, 146°00' W, 50-90 m, MNHN (2); DW 1952, 23°49' S, 147°53' W, 300-372 m, MNHN (1); DW 1 959, 23° 1 9' S, 1 49°30' W, 95-380 m, MNHN ( 1 ); DW 1996, 22°29' S, 15I°22' W, 489-1050 m, MNHN

(1) ; DW2013, 22°39' S, 152°50’ W, 80- 93 m, MNHN

(2) .

Distribution. Red Sea (type loeality), Madagascar, Maldives, Philippine Islands, New Caledonia, Solomon Islands, Fiji, French Polynesia (The Marquesas and the Austral Archipelago).

Remarks. Cytharomonila dollfusi was incorrectly synonymised with C. henedicta by Trôndlé & Houart (1992: 87, fig. 45). Mienis (2003) and Heiman and Mienis (2003) illustrated a specimen from the Gulf of Aqaba with its correct name.

See under C. paucimaculata for a comparison with that species.

Cytharomorula lefevreiana (Tapparone Canefri, 1 880 Text Fig. 1, Figs 5, 27-32

Tritonidea lefevreiana Tapparone Canefri, 1880: 65, pl. 3, figk 7-8.

Additional référencés

Pascula benedicta — Cemohorsky, 1982: fig. 20 only {noX Murex benedictus Melvill & Standen, 1895).

Monda squamilirata — Jairett, 2000: 60, fig. 247 (not Sistrum squamiliratum E.A. Smith, 1903).

NOT Pascula lefevreiana — Houart, 1995: 274, fig. 142; Tsuchiya, K. 2000: 385, fig. 105 (= Murex benedictus Melvill & Standen, 1895).

NOT Pascula lefevreiana - Houart, 2008; 204, pl. 397, figs la, \h {= Pascula ambonensis Wo\X2lx\, 1996).

NOT Cytharomorula lefevreiana — Houart, 2011: 280, fig. 6 {= Murex benedictus Melvill & Standen, 1895).

Type material. Two syntypes MZUF 24524, here selected as lectotype MZUF 24524/1 and paralectotypes MZUF 24524/2.

Type loeality. Mauritius.

Other material examined. Reunion. MD32, stn DC26, 21°22' S, 55°46' E, 310m, MNHN (1); stn DC41, 21°2r S, 55°27' E, 75m, MNHN (3); stn CP43, 21°21' S, 55°27'E, 73-77m, MNHN (1); stn DR47, 21°23'S, 55°36'E, 205-215 m, MNHN (4); stn DC56, 21°05'S, 55°12'E, 170-225m, MNHN (10); stn DC85, 20°59'S, 55°15'E, 58-70m, MNHN (47); stn DC86 20°59'S, 55°15'E, 75-90m, MNHN (1); stn CP97, 19°4rS, 54°09'E, 55m, MNHN (1); Reunion, Saline, 5- 20 m, MNHN (1); Reunion (no other data) (5), RH; Mauritius, ex coll. Jousseaume (5), MNHN.

29

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Classification of small species of Cytharomorula.

Distribution. Seychelles (Jarrett, 2000, as Monda scjuamilirata), Mauritius and Reunion.

Remarks. Cytharomorula lefevreiana differs from C. benedicta in being narrower and slenderer. The secondary and tertiary spiral cords are broader in C. lefevreiana and less numerous. The subsutural ramp is less concave with fewer and broader spiral sculpture, 3 or 4 cords in C. lefevreiana vs' 5 or 6 in C. benedicta.

There are also fewer spiral cords between PI and P2; 2 in C. lefevreiana 3 in C. benedicta, and between P2 and P3: 2 or 3 in P. lefevreiana vs 4-6 in C benedicta. The area between P3 and P6 is less concave than in C. benedicta.

Cytharomorula lefevreiana is correctly identified and iliustrated by Drivas & Jay (1988: 72, pl. 21, fig. 16). Another specimen was iliustrated as Pascula benedicta by Cemohorsky (1982) (see under C. benedicta).

Text Fig. 1. Original labels with

^ f -^y I r

^z''zz€y Z.4- Z ZI- -y . y^

ypé .

the syntypes of Tritonidea lefevreiana Tapparone Canefri, 1880 (photos Saulo Bambi)

Cytharomorula paucimaculata (Sowerby, 1903) Figs 10-13

Pentadactylus paucimaculatus Sowerby, 1903: 496.

Additional reference

NOT Cytharomorula paucimaculata — Houart, 2008: 198, pl. 394, figs 2-5; Houart & Héros, 2008: 459 (= Murex dollfusi Lamy, 1938).

Type material. Holotype NHMUK 1903.12.7.8 Type locality. Hachijojima (Hachijojima Island), Japan.

Other material examined. Japan, Okinawa, Seragaki, 19-25 m, under rocks, RH (2).

Distribution. Currently known from Japan in two localities: Seragaki (Okinawa) and Hachijojoma Island (east of Central Japan) (type locality).

Remarks. Cytharomorula paucimaculata differs from C. dollfusi in having a relatively broader shell with a broader, less elongate aperture, a comparatively shorter and broader siphonal canal and a higher spire.

The primary spiral cords are broader, almost of a same strength, while the P3 cord is usually weakly higher and stronger than Pl and P2 in C. dollfusi. The secondary and the tertiary cords are also broader and less numerous in C. paucimaculata.

The colour also differs, C. paucimaculata having a subsutural ramp with broad, light or dark brown coloured blotches, somewhat like in C. benedicta, and occasionally some brown dots on or between the primary spiral cords. C. dollfusi is usually unifomily light tan, rarely with some darker dots on and between the primary spiral cords.

The holotype of C. paucimaculata was iliustrated by Kaicher (1980: card 2542) as Evokesia paucimaculata and by Cemohorsky (1982: 129, figs 18-19) as Pascula benedicta forma paucimaculata. However, C. paucimaculata differs from C benedicta in having a comparatively larger shell with a higher spire, a broader aperture, a broader and shorter siphonal canal relative to the shell length, in having smaller and lower apertural denticles and in the P1-P4 spiral cords approximately similar in size, while the P3 cord is more obvions in C. benedicta and the P4 cord very small. The area between P3 and P6 is also much less concave than in C. benedicta.

Cytharomorula paucimaculata also has more numerous and lower axial ribs, 8-lOvs' 6 or 1 'm C. benedicta.

Figures 4-13

4. Spiral cord morphology of Cytharomorula ambonensis (Houart, 1996); 5. Spiral cord morphology of Cytharomorula lefevreiana (Tapparone Canefri, 1880); 6. Spiral cord morphology of Cytharomorula benedicta (Melvill & Standen, 1895).

7. Protoconch and first teleoconch whorl of Cytharomorula benedicta (Melvill & Standen, 1 895) (scale bar 500 pm). 8-9. Cytharomorula ambonensis (Houart, 1996). Indonesia, Ambon Id, Hitu, Suli, in sand, near coral, holotype RMNH 57155, 9.4 mm (scanned from a black and white photograph).

10- 13. Cytharomorula paucimaculata (Sowerby, 1903)

10. Hachijojima, Japan, holotype NHMUK 1903.12.7.8, 11.5 mm (scanned from a black and white photograph);

11- 13. Japan, Okinawa, Seragaki, 1 9-25 m, under rocks. 11-12. 12.4mm; 13. 11.5 mm.

30

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NOVAPEX 14(2); 25-34, 10 juin 2013

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Classification of small spccics of Cytharomonila.

Acknovvledgements

1 was able to examine the huge material of MNHN thanks to Philippe Bouchet (Muséum national d'Histoire naturelle, Paris) who permits me to study the Muricidae collected during the expéditions conducted by MNHN and IRD in the Tropical Indo- Pacific and to Virginie Héros who helped me to search the Cytharomonila and Pascula speeimens in their collections. Thanks also to Virginie Héros and Philippe Maestrati for the digital photographs of Murex dollfusi.

Thanks also to Shawn Miller, Okinawa, Japan, for the two speeimens of Cytharomonila paiicimaciilata he kindly sent to me.

The photo of the holotype of C. paiicimaciilata was taken during my visit to the National Muséum of Natural History, in the late eighties.

I was able to illustrate the type material of Murex benedictus thanks to Henry Mcghie and Rebecca Machin, The Manchester Muséum, UK, who provided the digital photos of the figured syntype.

The search for the type material of Tritonidae lefevreiana lasted several months and ended suceessfully thanks to Philippe Bouehet, Muséum national d'Histoire naturelle, Paris, France; for useful information, to Marco Oliverio, Universita di Roma "La Sapienza" Dipartimento di Biologia Animale e dell'Uomo, Roma, Italy; to Giuliano Doria from the Museo Civico di Storia Naturale "Giacomo Doria", Genova, Italy, and to Simone Cianfanelli, Museo di Storia Naturale, Università di Firenze, Italy for also giving me useful infonnation about possible contacts; and fmally, thanks to the kindness of Cecilia Volpi and Antonio Callea, Museo di Storia Naturale, Sezione di Zoologia "La Specola", Università di Firenze, Italy, for searching and finding these types and to Saulo Bambi for the very fine digital photographs.

I am also very grateful for the continuing help of Anders Warén, Natural History Muséum, Stockholm, Sweden, in the préparation and SEM work of the radula.

I also really appreciate the help of John Wolff, Lancaster, Pennsylvania, USA, who regularly checks and comects my English and brings other comments to my papers, including this one.

Finally, thanks to an unknown referee for some linguistic suggestions

REFERENCES

Cemohorsky, W.O. 1982. The taxonomy of some Indo-Pacific Mollusca, part 10, Records ofthe Auckland Institute and Muséum 19; 125-147. Claremont M., Houart, R., Williams S. T. & Reid D.G. 2013. A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae).

Journal of MoUiiscan Stiidies, 79: 19-29.

Drivas, J. & Jay, M. 1988. Coquillages de la Réunion et de nie Maurice. Delachaux & Niestlé, Neuchâtel-Paris: 1-159.

Heiman, E.E. & Mienis, H. K. 2003. Shell of East Sinai, an illustrated list, Muricidae (2). Triton 8: 22-24.

Higo, S., Callomon, R, & Goto, Y. 1999. Catalogue and bibliography of the marine shell-bearing Mollusca of Japan. Gastropoda. Bivalvia. Polyplacophora. Scaphopoda. Elle Scientific Publications, Japan, 749 pp.

Houart, R., 1995 ("1994"). The Ergalataxinae

(Gastropoda, Muricidae) from the New Caledonia région with some comments on the subfamily and the description of thirteen new species from the Indo-West Pacific. Bulletin du Muséum national d' Histoire naturelle, Paris, 4e sér., 16, section A, n° 2-4: 245-197.

Figures 14-40

14-19. Cytharomonila ambonensis (Houart, 1996)

14-15. Guam, Apra Harbor, among silty rocks; glass breakwater, 5-9 m, RH, 10.1 mm; 16-17. Guam, near mouth of Apra Harbor, among rocks; glass breakwater, 5-8 m, RH, 8.8 mm; 18-19. Lifou, New Caledonia, MNHN IM- 2010-21107, 6.3 mm.

20-26. Cytharomonila benedicta (Melvill & Standen, 1895)

20-21. Lifou, New Caledonia, Lectotype Manchester Mus. EE.3708, 6.7 mm; 22-23. Koumac, New Caledonia, RH, 7.4 mm; 24-25. Koumac, 15-20 m, RH, 6.7 mm; 26. New Caledonia, Loyalty Islands, Lifou, 20°48' S,

1 67°07' E, 8- 1 8 m, MNHN IM-20 10-21118, 7.4 mm.

27-32, Cytharomonila lefevreiana (Tapparone Canefri, 1880)

27-28. Mauritius, Lectotype MZUF 24524/1, 1 1 mm; 29. Paralectotype MZUF 24524/2, 8.3 mm; 30-32. Reunion, 21°00' S, 55° 15' E, 58-70 m, MNHN. 30-31. 6.6 mm; 32. 7 mm.

33-40. Cytharomonila dollfusi (Lamy, 1938)

33-34. Jubal Island, holotype MNHN 0103, 8 mm; 35-36. Philippines, Cebu, Mactan Id, Punta Engano, RH, 8.5 mm; 37. Phillippines, Sulu Sea, RH, 10.4 mm; 38. Maldives, Maagau Kandu, Ari Atoll, 20-25 m, RH, 9.8 mm; 39- 40. New Caledonia, Eoyalty Islands, Santal Bay, 20°48' S, 167°08' E, 35-45 m, MNHN lM-20 10-21 111, 10.9 mm.

32

R. Houart

NOVAPEX 14(2); 25-34, lO jLiin 2013

33

R. H ou A RT

Classification of small species of Cytharomonila.

Houart, R. 1996. Results of the Rumphius

Biohistorical Expédition to Ambon (1990). Part. 5. Mollusca, Gastropoda, Muricidae. Zoologische Mededelingen 70: 377-397.

Houart, R. 2008. Muricidae. In: Poppe G. (ed.), Philippine Marine Molliisks II, Conehbooks, Haekenheim, Germany: 132-220.

Houart, R. 2011. Family Muricidae -Review and eomments. In Sevems, M. Shells ofthe Hawaiian Islands - The Sea Shells: 280-294, pis 123-130. Conehbooks, Haekenheim, Gennany: 1-564. Houart, R. & Gofas, S. 2012. Cytharomonila Kuroda, 1953. Aecessed through: World Register of Marine Speeies at

http://www.marinespeeies.org/aphia.php7pMaxdet ails&id=138190 on 28 Nov. 2012.

Houart, R. & Héros, V. 2008. Muricidae (Mollusca: Gastropoda) from Fiji and Tonga, in Héros V, Cowie R. H. & Bouchet, P. (eds), Tropical Deep- Sea Benthos 25. Mémoires du Muséum national d'Histoire naturelle 196: 437-480.

Houart, R. & Trôndlé, J. 2008. Update of Muricidae (excluding Coralliophilinae) from French Polynesia with description of ten new species. Novapex 9(2-3): 53-93.

Jarrett, A. G. 2000. Marine shells of the Seychelles. Carole Green Publishing, Cambridge, U. K.: i-xiv, 1-149.

Kaieher, S.D. 1980. Card catalogue of world-wide shells, Muricidae V. Privately publ. St. Petersburg, Florida.

Famy, F. 1938. Mission Robert Ph. Dollfus en Egypte: VII. Mollusca Testacea. Mémoires de l'Institut d'Egypte 37: 1-90.

Melvill, J.C. & Standen, R. 1895. Notes on a

collection of shells from Fifu and Uvea, Foyalty Islands fomied by the Rev. James and Mrs. Hadfield, with list of speeies. Journal of Conchology 8: 84-132.

Merle, D. 1999. La radiation des Muricidae {Gastropoda : Neogastropoda) au Paléogène: approche phylogénétique et évolutive. Paris. Unpublished thesis, Muséum national d'Histoire naturelle : i-vi, 499 pp.

Merle, D. 2001. The spiral cords and the internai dentieles of the outer lip in the Muricidae: temiinology and methodological eomments. Novapex 2(3): 69-91.

Mienis, H. K. 2003. Notes on some muricid gastropods from the Gulf of Aqaba. Triton 8: 7-8.

Sowerby, G. B. 1903. Descriptions of fourteen new species of marine Mollusca from Japan. The Aimais and magazine ofNatural Histoiy. Vol. 12(7): 496-501.

Tapparone Canefri, C.M. 1880. Glanures dans la faune malacologique de l'Ile Maurice - catalogue de la famille des Muricidés (Woodward).

Mémoires de la Société Malacalogique de Belgique 15: 1-103.

Trôndlé, J. & Houart, R. 1992. Fes Muricidae de Polynésie Française. Apex 1: 67-149.

Tsuchiya, K. 2000. Muricidae. In: Okutani, T. (ed.). Marine Mollusks in Japan, Tokai University Press, Tokyo: 364-421.

34

R. Goethals & D. Monsecour

NOVAPEX 14(2): 35-37, 10 juin 2013

A new species of Rolaniconus (Gastropoda: Conidae) from the Philippines

Rika GOETHAELS

Melsbroeksestraat 2 1 , 1800 Peutie-Vilvoorde, Belgium fernand.de.donder@pandora.be

David MONSECOUR Dahliastraat 24, 3200 Aarschot, Belgium david.monsecour@telenet.be

Keywords. Gastropoda, Conidae, Conus, Rolaniconus dedontleri new speeies, Philippines.

Abstract. Rolaniconus dedonderi is described as a new species and compared with Conus axelrodi (Walls, 1978) and Rolaniconus lecourtorum Lorenz, 2011. The new species has hitherto been confused with R. axelrodi and has consistently been identified as an albino form of this species.

INTRODUCTION

While sorting hundreds of smaller Conidae originating from the Palawan area, southem Philippines, the senior author encountered a number of specimens that were at first identified as Rolaniconus axelrodi (Walls, 1978). Yet, a more thorough survey of the original description and figures of the types (Walls, 1978 and 1979; Kohn & Anderson) revealed a number of constant characteristics not encountered in R. axelrodi and led to the conclusion that the specimens at hand belonged to a yet undescribed species. They are described here as Rolaniconus dedonderi sp. nov.

The terminology used here follows Rôckel & al. (1995)

Abbreviations

MNHN: Muséum national d’Histoire naturelle, Paris, France.

DG: Collection Fernand & Rika De Donder - Goethaels, Peutie, Belgium.

DM: Collection David Monsecour, Aarschot,

Belgium.

SYSTEMATICS

Family CONIDAE Fleming, 1822 Genus Rolaniconus Tucker & Tenorio, 2009 Type species: Conus varius Linnaeus, 1758 by original désignation.

Rolaniconus dedonderi sp. nov.

Figs 1-5

Type material. Flolotype MNHN 25879, 20.8 x 11.5 mm. Paratypes 1-3 (from type locality) DG, 20.1 - 22.3 mm, Paratypes 4-5 (Balabac Island, Palawan, southem Philippines) DM, 17.4 and 22.7 mm.

Type locality. Coron, Palawan Island, southem Philippines, 20-35m.

Distribution and habitat. Ail available material originates from the Palawan area, southem Philippines. The distributional range is probably limited to these régions, but could be much larger because of the confusion with R. axelrodi (see below). No real data about the habitat are available at présent, except for a bathymetrical range of 20-35 métrés.

Description. Shell of rather small size for the genus (about 17-23 mm in length), ventricosely conical, outline slightly sigmoid; shoulder subangulate, undulate; spire slightly concave. Protoconch of about 2 bulbous whorls adomed with microscopie pits, glassy but not translucent (eroded in holotype and most of the paratypes). Teleoconch consisting of 8 whorls with 4 evenly spaced spiral cords on ail whorls; shoulders adomed with low, rounded knobs. Spiral sculpture of last whorl consisting of 8-10 cords on abapical 1/3, lower and less distinct adapically, obsolète or microscopie from final visible cord towards shoulder. Visible cords continuous, sometimes with rounded, elongated élévations (only visible under magnification). Lip simple, aperture of nearly uniform width. Outer apertural lip completely smooth. Columella like last whorl: abapical spiral cords continue and run deep into the aperture, remaining part of columella virtually smooth, occasionally with microscopie cords.

Colour of protoconch glassy off-white, sometimes with pale yellowish tinge on it. Rest of shell unifonnly white.

Comparison. Rolaniconus dedonderi has hitherto always been considered a white or even albino fomi of R. axelrodi. White specimens of the latter are known (Raybaudi Massillia: pl. 567, fig. 3), but the new species can easily be distinguished by the weaker spiral sculpture on the last whorl: obsolète or

35

R. GOETllALS & D. MONSECOUR

A new species of Rolaniconiis.

microscopie between the shoulder and the basal cord, whereas the last whorl of R. axelrodi is completely adorned with strong spiral cords of beadlike nodules; the higher onset of the lip at the shoulder, whereas it is Lisually somewhat lower in R. axelrodi; the “straighter” general outline of the body whorl; the less bulbous teleoconch whorls and the pure white colour. Even thoLigh both species occur in the southem Philippine région around Palawan, R. dedonderi lives in deeper waters (20-35 m as opposed to 8-15 m for R. axelrodi), but more accurate data are needed to confimi this.

In fact, R. dedonderi cannot be confused with any other known species, but as Lorenz (201 1) compared R. lecourtorum Lorenz, 2011 with A. axelrodi in his original description, we hereby also compare R. dedonderi and R. lecourtorum: R. dedonderi has a weaker spiral sculpture, which does not cover the entire last whorl, the shoulders bear low, rounded knobs and it is completely white, whereas R. lecourtorum has a stronger, nodulose spiral sculpture which is présent ail over the last whorl, a stepped spire with strong coronations and a reddish colour pattern. Moreover, R. dedonderi is only known from the Philippines, whereas R. lecourtorum is endemic to the Cargados Carajos Shoals (also known as St. Brandon) in the western Indian Océan.

Etymology. The new species is named in honour of Lemand De Donder, the first author’s husband, for his

efforts to draw attention to the smaller, often neglected molluscan species from the Philippines.

Acknowledgements

We would like to thank Mike Lilmer for his opinion on R. dedonderi and Kevin Monsecour for making the photographs and composing the plate.

REFERENCES

Kohn, A. J. & Anderson, T. The Conus biodiversity website:

http://biology.burke.washington.edu/conus (last accessed 20/07/2012).

Lorenz, P. , 201 1 . A new species of Rolaniconus from the western Indian Océan (Gastropoda : Conidae). Schriften zur Malakozoologie ans dem Haus der Natur— Cismar 26: 37-40.

Raybaudi Massilia, G., 2008. Conidae. In: Poppe,

G. T. Philippine Marine Mollusks. Vol. 2. ConchBooks, Hackenheim.

Rôckel, D., Kom, W. & Kohn, A. J., 1995. Manual of the Living Conidae. Volume 1: Indo-Pacific Région. Verlag Christa Hemmen, Wiesbaden: 1- 517.

Walls, J. G. 1978. Diagnoses of four new cônes (Mollusca: Conidae). The Pariah 2: 1-7.

Walls, J. G., 1979. Cône shells. A synopsis of the living Conidae . T. F. H. Publications, Neptune City, New Jersey. 1-1011.

Figures 1-8

1-5, Rolaniconus dedonderi sp. nov.

1-2. Holotype MNHN 25879, Coron, Palawan Island, Southern Philippines. 20-30m deep, 20.8 x 1 1.5 mm;

3. Paratype 1. Type locality, 20.0 mm, coll. DG; 4-5. Paratype 2. Type locality, 19.0mm, coll. DG.

6-8, Rolaniconus axelrodi (Walls, 1978)

Coron, Palawan Island, Southern Philippines. 20-30m deep (Figs. 6-7. 18.8. mm; Fig. 8. 16.9 mm), both coll. DM.

36

R. Goethals & D. Monshcour

NOVAREX 14(2): 35-37, 10 juin 2013

37

K. Fraussen & P. Stahlschmidt

NOVAPEX 14(2): 39-41, 10 juin 2013

Eclectofusus^ a new genus for Pararetifusus dedonderi Fraussen & Hadorn, 2006 (Gastropoda: Buccinidae)

Koen FRAUSSEN

Leuvensestraat 25, B-3200 Aarschot, Belgium koen.fraLissen@skynet.be

Peter STAHLSCHMIDT Institute for Environmental Sciences Universitàt Koblenz-Landau Fortstrasse 7, D-76829 Landau, Germany stahlschmidt@uni-landau.de

Keywords. Gastropoda, Buccinidae, Pararetifusus, Indo-Pacific, new genus.

Abstract. Eclectofusus gen. nov. (Bueeinidae) is described to accomodate Pararetifusus dedonderi Fraussen & Hadom, 2001. The new genus is eompared with Pararetifusus Kosuge, 1967 and Americominella Klappenbaeh & Ureta, 1972.

INTRODUCTION

The deseription of Pararetifusus dedonderi Fraussen & Hadom, 2001 was not aeeompagnied by the deseription of a new genus to prevent the establishment of a monotypic genus. That prineiple worked well until a révision of Pararetifusus was published by Kosyan (2006). In the meantime we searehed through material of many expéditions and muséums to fmd more speeies whieh may belong to this group but without resuit. In the présent paper we deseribe the new genus Eclectofusus to aeeommodate "'Pararetifusus'" dedonderi properly. Previously only known from the Philippines, the range of E. dedonderi is hereby extended to Vanuatu and the Solomon Islands.

Bathymétrie distributions, previously only based on information obtained from fishennen and shell dealers, are here based on detailed information from the seientifie expéditions.

Material and methods

The material reported in the présent study originates from varions expéditions conducted sinee 1980 by MNHN and IRD (formerly ORSTOM):

(a) SALOMON 1 (Solomon Islands).

(b) MUSORSTOM 3 (northern Philippines).

(e) PANGLAO 2005 (Bohol and Sulu Seas;

Philippines).

(d) MUSORSTOM 8 (Vanuatu).

Material from these expéditions is, unless otherwise stated, deposited in MNHN.

We provided the bathymétrie range whieh is reported as the broadest possible depth range and the so called eonfimied depth range whieh is the nairowest possible range for E. dedonderi (e.g. if a speeimen was

reported from a depth of 100-200 m and a seeond speeimen of the same speeies from 500-600 m, the bathymetie range would be 100-600 m, while the eonfiiTned bathymétrie range would be 200-500 m). The bathymétrie range takes into aeeount records of dead and live taken specimens while the eonfirmed depth range is only provided for live taken specimens. The depth ranges of every station were assigned to ‘200 m’ categories (e.g. 0-200 m, 200-400m, ...). If a depth range was not unequivoeally assignable, the eategory eomprising the corresponding mid point of the depth range was used (e.g. a depth range of 320- 440 m with the mid point at 380 m would beeome assigned to the ‘200-400 m’ eategory). See Fig. 1.

Abbreviations

CP: Chalut à perehe (beam trawl).

DW: Drague Warén (Warén dredge).

IRD: Institut de Reeherche pour le Développement (Fomierly ORSTOM).

KBIN: Koninklijk Belgiseh Instituut voor Natuurwetensehappen, Brussels, Belgium.

KF: eollection Koen Fraussen, Aarsehot. Belgium. MNHN: Muséum national d'Histoire naturelle, Paris, Franee.

ORSTOM: Office de la Recherche Seientifique et Teehnique d'Outre Mer (now IRD), dd: empty shell, dead colleeted.

Iv: eollected alive.

jv: juvénile or subadult speeimen/shell. SYSTEMATICS

Family BUCCINIDAE Rafmesque, 1815 Eclectofusus gen. nov.

39

K. Frausscn & P. Stahlschmidt

Eclectofusus gen. nov.

Type species. Pararetifusiis dedonderi Fraussen & Fladom, 2001 (type locality: Balicasag Island, near Panglao, Bohol, central Philippine Islands, in 120-250 m).

Range. The single species, Eclectofusus dedonderi comb. nov., is known from the Philippines, the Solomon Islands and Vanuatu.

Diagnosis. Shell small, up to 19.4 mm in length, usually between 12 and 17 mm, thin, rather fragile, semi-transparant. Shape broadly fusifomr, spire high. Siphonal canal short, narrow, open.

Protoconch paucispiral, glossy, smooth.

Sculpture reticulate, consisting of fine, sharp spiral cords Crossing nan'ow but prominant axial ribs, fonning a sharp knob at intersections.

Aperture oval, outer lip usually thin (subadult), occasionally thick, with 2-7 big denticles within (adult specimens), columella smooth without columellar knob, glossy, slightly curved. Operculum slightly smaller than aperture, comeous, pale brown; shape rather triangular, pointed, nucléus tenninal. Periostracum thick, rather comeous, dark brown; fonning flamboyent sharp axial lamellae with spiny hairs on intersections of spiral cords with axial ribs, these spiny hairs occasionally bicuspid on shoulder. Radula with tricuspid outer teeth.

Comparison, Eclectofusus gen. nov. is characterized by having a reticulate sculpture in combination with big and rounded internai denticles inside the outer lip, a hirsute periostracum and a small adult size. Pciraretifusus Kosuge, 1967 (type species: Phymorhynchus tennis Okutani, 1966: 26, Japan) differs in having two or three pronounced spiral cords and by the absence of axial sculpture.

Americominella Klappenbach & Ureta, 1972 [type species: Americominella duartei Klappenbach & Ureta, 1972: 2, Argentina (= A. longisetosus De Castellanos & Fernandez, 1972, Argentina)] ( = Echinosipho Kaiser, 1977 with type species: Echinosipho aculeatum Kaiser, 1977: 27, Argentina, a synonym of A. duartei) is similar in sculpture and periostracum. The radula is quite identical with the exception of the central tooth which has a concave base in A. duartei (instead of being rectangular) and the broader base of the latéral teeth (Bouchet & Warén, 1986: pl. 2, fig. 10). Furthermore, A. duartei differs at once in having a broad, depressed protoconch consisting of 1.5 whorls, a larger size (up to 90 mm) and a different distribution (subantarctic, off Argentina).

Etymology. Eclectofusus gen. nov. is named after eclectisism, one of the artistic trends during the romanticism (1820-1870) which is based on compilating distinct styles, referring to the characteristics of this shell which one may regard as a compilation from a varicd number of families: the

shape of Eiisinus (Fasciolariidae), the usual colour and also the sculpture of Trophon (Muricidae), the reticulate sculpture of Raphitoma (Raphitomidae), the periostracum of Cymatium (Ranellidae) but the radula of Buccinidae.

Eclectofusus dedonderi (Fraussen & Hadom, 2001) comb. nov.

Fig. 2

Pararetifusus dedonderi Fraussen & Hadom, 2001: 93.

Type locality. Balicasag Island, near Panglao, Bohol, central Philippines, in 120-250 m.

Type material. Holotype in KBIN 490.I.G.nr.29.260 and 1 8 paratypes.

Other material examined. A total of 15 lots (23 specimens). Philippines: MUSORSTOM 3: Stn CP139, 11°53’N, 122°14’E, 240-267 m, 1 Iv, PANGLAO 2005 (Bohol Sea), stn CP2343 (Pamilacan Island), 9°27’N, 123°49’E, 273-356 m, 1 dd, stn DW2364 (Dipolog), 9°01’N, 123°25’E, 427 m, 2 dd (1 jv), stn CP2380 (Dipolog), 8°41’N, 123°18’E, 150-163 m, 4 Iv (1 jv), stn CP2383 (Aliguay Island), 8°45’N, 123°18’E, 338-351 m, 1 Iv,

1 dd (1 Jv), stn CP2384 (Aliguay Island), 8°46’N, 123°16’E, 624-647 m, 1 Iv, 1 dd, stn CP2392 (Balicasag Island), 9°29’N, 123°41’E, 242-400 m, 1 Iv, 1 dd, stn CP2393 (Balicasag Island), 9°30’N, 123°42’E, 356-396 m, 1 Iv, stn CP2404 (Maribojoc Bay), 9°39’N, 123°43’E, 481-505 m, 1 Iv, stn CP2409 (Maribojoc Bay), 9°45’N, 123°45’E, 220- 257 m, 1 Iv.

Solomon Islands: SALOMON 1, stn CP 1804, 9°32’S, 160°37’E, 309-328 m, 1 Iv (jv), stn CP1831, 10°12’S, 161°19’E, 135-325 m, 2 Iv (1 jv), stn CP1837, 10°13’S, 161°29’E, 381-383 m, 1 dd, stn CP 1860, 9°22’S, 160°31’E, 620 m, wood, 1 Iv. Vanuatu: MUSORSTOM 8, stn DW978, 19°23’S, 169°27’E, 408-413 m, 1 dd.

Range and habitat. Western Pacific, before only reported from the central Philippines, the range is extended towards the south-east to the Solomon Islands and Vanuatu. The confirmed bathymétrie range is 163 to 624 m for live collected specimens, records of empty shells range from 135 to 647 m.

Live specimens were sampled on sandy-muddy bottoms with volcanic mud and/or coral rubble.

The species is syntopic with Manaria species at the Philippines (PANGLAO 2005, stns CP2343, CP2384, CP2393) and at the Solomon Islands (SALOMON 1, stns CP 1804, CP 1837, CP 1860) ) and with Preangeria dentaia at the Philippines (PANGLAO 2005, stns CP2380, CP2383,CP2392).

40

K. Fraussen & P. Stahlschmidt

Novapex 14(2): 39-41, 10 juin 2013

Number of specimens

Figure 1 Bathymétrie distribution of MNHN material of Eclectofiisiis dedonderi.

Reniarks. Eclectofusus dedonderi comb. nov. is characterized by a strong, reticulate sculpture in combination with a small, smooth, paucispiral protoconch and a reticulate and hirsute periostracum.

Figure 2. Eclectofusus dedonderi comb. nov., 11.6 mm, paratype 12, Philippines, Balicasag Island, in tangle nets, 120-150 m deep, KF-3206.

Genus Pararetifmus Kosuge, 1 967

Pararetifusus Kosuge, 1967: 59-64.

Type species: Phymorhynchus tennis Okutani, 1966: 26, pl. 11-21 (type locality: Japan, Sagami Bay, 34°57’N, 139°2rE, 1470-1500 m).

Remarks. Pararetifusus species are mainly known from the Boréal and Arctic région. Three species are known. For a discussion of included species we refer to Kosyan, 2006 (5-15).

Eclectofusus sp. nov. differs in having sharp axial ribs, fonning a reticulate sculpture together with the spiral sculpture.

Included species

Pararetifusus kantori Kosyan, 2006 (type locality: Russia, Bering Sea, 52°40.8’N, 159°13’E, 800-1000 m).

Pararetifusus kosugei Kosyan, 2006 (type locality: Russia, Bering Island, 55°08’N, 165°58.3’E, 130-250 m).

Pararetifusus tennis (Okutani, 1966) (type locality: Japan, Sagami Bay, 34°57’N, 139°2rE., 1470-1500 m).

Acknowledgments

We are grateful to Philippe Bouchet, Virginie Héros and Philippe Maestrati (Muséum national d'Histoire naturelle, Paris, France) for procuring additional material for study, to Roland Houart (Belgium) for reading and correcting the manuscript and to David Monsecour (Belgium) for correcting the English text.

REFERENCES

Bouchet, Ph. & Warén, A. 1986. Mollusca

Gastropoda: Taxonomical notes on tropical deep water Buccinidae with descriptions of new taxa, in: Résultats des Campagnes MUSORSTOM. 1 & II. Philippines (1976-1980). Tome 2. Mémoires de Muséum national d’Histoire naturelle, Paris, série A, Zoology 133: 455-499, text figs.1-4, pis. 1-18. Castellanos, Z. J. A. de & Fernandez, D. 1972. Un nuevo Bathydomus para aguas subantarcticas (Mollusca, Buccinulidae). Neotropica, 18: 81-86. Fraussen, K. & Hadom, R. 2001. A new species of Buccinidae from the Philippine Islands. Novapex, 2(3): 93-96.

Kaiser, P. 1977. Über den Fund einer neuen

Buccinidae (Mollusca), Echinosipho aculeatum gen. n. und sp. n. in patagonischen Gewâssem. Mitteilungen ans den Hamburgischen Zoologischen Muséum und Instituut, 74: 27-30. Klappenbach, M. A. & Ureta, E. H. 1972. Nuevo

genero y nuevo especie de la familia Buccinidae de aguas Uruguayas y Argentinas. Comunicaciones Zoolôgicas del Museo de Historia Natural de Montevideo, 10(134): 1-6.

Kosuge, T. 1967. On the transfer of" Phymorhynchus ?" tennis Okutani, 1966 to the family Buccinidae. Venus, 25: 59-64.

Kosyan, A. R. 2006. Two new species of the genus Parretifusus Kosuge, 1967 (Buccinidae: Colinae), with notes on the moiphology of Pararetifusus tennis (Okutani, 1966). Ruthenica, 16(1-2): 5-15. Okutani, T. 1966. Archibenthal and abyssal Mollusca collected by the R. V. Soya-Maru from Japanese waters during 1964. Bulletin ofthe Tokai Regonal Eisheries Laboratoiy, 46: 1-35.

41

%

R. Houart & M. Severns

Novapex 14(2); 43-48, lOjuin 2013

Description of a new species of Favartia (Pygmaepterys) (Gastropoda: Muricidae: Muricopsinae) from Hawaii

Roland HOUART

Research Associate, Institut royal des Sciences naturelles de Belgique Rue Vautier, 29, B- 1000 Bruxelles, Belgium roland. houart (^skynet.be

Mike SEVERNS 3415 Kehala Drive Kihei, Hawaii, USA 96753 msevems@hawaii.rr.com

Keywords. Gastropoda, Muricidae, Muricopsinae, Hawaii, Favartia (Pygmaepteiys) new species.

Abstract. Favartia {Pygmaepterys) kernoi n. sp. is described from Hawaii. It was previousiy confused with F. {P.) funafutiensis (Hedley, 1899) but differs in having a lecithotrophic larval development vs planktotrophic in F. {P.) funafutiensis. The new species is compared with other Indo-West Pacific species of Pygmaepteiys, namely F. (P.) avatea Houart & Trôndlé, 2008 from French Polynesia and New Caledonia, F. {P.) bellini (D'Attilio & Myers, 1985) from the Philippines and Okinawa, F. {P.) lifoiiensis Houart & Héros, 2012 from New Caledonia and F. {P.) philcloveri (Houart, 1984) from the Philippines. It is also briefly compared with the other eleven species from the Indo-West Pacific.

Résumé. Favartia {Pygmaepterys) kernoi n. sp. est décrite de Hawaii. L'espèce a été confondue avec F. {P.) funafutiensis (Hedley, 1899) mais elle en diffère par son développement larvaire lécithotrophe alors qu'il est planctotrophe chez F. {P.) funafutiensis. La nouvelle espèce est comparée avec d'autres espèces de Pygmaepteiys de l'Indo-Pacifique Ouest, notamment F. {P.) avatea Houart & Trôndlé, 2008 de Polynésie française et de Nouvelle-Calédonie, F. {P.) bellini (D'Attilio & Myers, 1985) des Philippines et d'Okinawa, F. {P.) lifoiiensis Houart & Héros, 2012 de Nouvelle-Calédonie et F. {P.) philcloveri (Houart, 1984) des Philippines. Elle est également brièvement comparée avec les onze autres espèces de l'Indo-Pacifique Ouest.

INTRODUCTION

Pygmaepterys was originally described as a subgenus of Pterynotus s. s. for two species: F. {P.) alfredensis (Bartsch, 1915), the type species, and F. {P.) maraisi (Vokes, 1978), both from South Africa. It was then thought that Pygmaepteiys was closely related to Pterynotus. However, further research based on Shell morphology and radula characteristics proved Pygmaepterys to be a muricopsine (D'Attilio & Myers, 1985b). The radula characteristics were studied in several species but not in the type species; nevertheless, ail these species are strongly related to F. alfredensis.

There are currently 24 Recent Pygmaepteiys species, 16 of which are from the Indo-West Pacific région. The new species is the only one that occurs in Hawaii.

Terminology used to describe the spiral cords and the apertural denticles (after Merle, 2001 and 2005) (Figs 1-2) (Terminology in parenthèses: erratic feature).

SPIRAL CORDS

P: primary cord; s: secondary cord; IP: infrasutural primary cord (primary cord on subsutural ramp); PI:

shoulder cord; P2-P6: primary cords of the convex part of the teleoconch whorl; sl-s6: secondary cords of the convex part of the teleoconch whorl (example; si = secondary cord between PI and P2; s2 = secondaiy cord between P2 and P3, etc.); ADP: adapertural primary cord on the siphonal canal; MP: médian primaiy cord on the siphonal canal; ABP: abapertural primary cord on the siphonal canal.

APERTURE

ID: infrasutural denticle; DI to D5: abapical denticles.

Repositories

AMS: Australian Muséum, Sydney, Australia.

ANSP: Academy ofNatural Sciences of Philadelphia, U.S.A.

IRSNB: Institut royal des Sciences naturelles de Belgique, Bruxelles, Belgium.

MNHN; Muséum national d'Histoire naturelle, Paris, France.

MS: collection Mike Severns.

RH: collection Roland Houart.

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New species of Favartiu (Pyginaeplerys) from Hawaii.

SDNHM: San Diego Natural Histoiy Muséum, California, U.S.A.

USNM; National Muséum of Natural History, Washington, D.C., U.S.A.

Other abbreviations

Iv: live collected specimen. dd: empty shell.

SYSTEMATICS

Family MURICIDAE Rafinesque, 1815

Subfainily MURICOPSINAE Radwin & D'Attilio, 1971

Genus Favartia Jousseaume, 1880

Subgenus Pygmaepterys Vokes, 1978

Type species by original désignation: Murex

alfredensis Bartsch, 1915, South Africa.

Remarks. In hannony with the most recent papers (Houart & Trôndlé, 2008; Houart & Rosado, 2008; Houart & Héros, 2012) we will retain Pygmaepteiys as a subgenus of Favartia, while awaiting further genetic results.

Favartia {Pygmaepterys) kernoi n.sp.

Figs 1-4, 6, 9-14

Favartia funafutiensis (Hedley) - Kaicher, 1979: card 2029 (NOT Murex funafutiensis Hedley, 1899). Pygmaepterys funafutiensis (Hedley) - Vokes and D'Attilio, 1980: PI. 2, fig. 5a-b (only) (NOT Murex funafutiensis Hedley, 1899).

Pygmaepterys funafutiensis (Hedley) - D'Attilio & Myers, 1985a: Figs 7-8, 11, 14; D'Attilio & Myers, 1985b: Figs 4, 5 (NOT Murex funafutiensis Hedley, 1899).

Type material. Hawaii, dredged off Kihei, Maui, 20°39.180" N, 156°28.598" W to 20°38.650" N, 156°30.494" W, 170 m, December 28, 2012, holotype MNHN 25838, Iv, 10.7 mm.

Paratypes : 1 IRSNB IG 32 388/MT 2798, Iv, 9.5 mm; 1 RH, dd, 10.5 mm; 1 MS, dd, 11.8 mm; 1 ANSP 451390, dd, 9.9 mm; 1 MS, dd, 8.7, 128 m, November 21, 2012; 1 USNM 1205569., Iv, 9.2 mm, 128 m, November 21, 2012; 1 MS, Iv, 13.3 mm; 137 m, November 30, 2012; 1 MS, dd, 12.9 mm, 101 m, December 5, 2012; Hawaii, Oahu, Pokai Bay, August 14, 1976, Dredged in sand & coralline algae, 183 m, 1 R. Salisbury, dd, 9.0 mm.

Type locality. Hawaii, dredged off Kihei, Maui, 20°39.180" N, 156°28.598" W to 20°38.650" N, 156°30.494" W, 170 m, December 28, 2012.

Distribution. Hawaii, Maui and Oahu, live in 101- 1 70 m.

Description. Shell large for the subgenus, up to 13.3 mm in length at maturity (paratype MS). Length/width ratio 1.7- 1.8. Biconical, broadly ovate, weakly spinose, lightly built, squamous. Subsutural ramp narrow, weakly sloping, slightly concave. Light or dark tan or brown. Aperture glossy, bluish-white.

Spire high with 1.5 protoconch whorls and teleoconch up to 5 broadly convex, strongly shouldered, weakly spinose whorls. Suture slightly impressed, partially obscured by axial varices of following whorl. Protoconch moderately large, weakly elongate, whorls rounded, smooth, width 550 pm, height 500 pm (paratype IRSNB). Temiinal lip délicate, thin, erect, strongly curved.

Axial sculpture of teleoconch whorls consisting of moderately high, narrow, weakly spinose, lightly webbed varices. Other axial sculpture of very fine, numerous, close-set growth lamellae. First to penultimate whorl with 6 varices; last whorl with 5 or 6. Spiral sculpture of high, strong, rounded, broad, squamous, primary cords and several lirae. First to third whorl with visible PI and P2, fourth with SP, PI, P2, occasionally P3 covered by next whorl. Last whorl with SP, P1-P6, with small lirae between cords. P2-P4 broadest and highest, approximately of same strength; PI and P5 slightly lower and narrower; P6 very small, followed by ADP and MP on siphonal canal. Intersection of varices and spiral cords giving rise to small, short spinelets, connected with varical flange, more obvions on last (apertural) varix. PI spine longest.

Aperture small, ovate; columellar lip naiTOw, with 3 weak knobs abapically, otheiAvise smooth; rim veiy weakly erect abapically, otheiAvise adhèrent, with weak, low pariétal tooth at adapical extremity; anal notch moderately deep, broad; outer lip erect, with 6 strong denticles within: ID, D1-D5. Siphonal canal moderately long, 28-30% of total shell length, broad, weakly dorsally reciuwed, narrowly open.

Operculum and radula unknown.

Figures 1-8 (scale bars: 500 pm)

1-4. Favartia {Pygmaepteiys) kernoi n. sp.

1. Apertural denticles morphology (holotype MNHN 25838.); 2. Spiral cords morphology (paratype RH); 3. Measurements of the shell and siphonal canal (holotype MNHN 25838.); 4. Protoconch (paratype IRSNB IG 32388/MT 2798).

5. Protoconch of Favartia {Pygmaepteiys) funafutiensis (Hedley, 1899), holotype AMS C. 006004.

6-8. Morphology of the aperture.

6. Favartia {Pygmaepterys) kernoi n. sp. (holotype MNHN 25838); 7. Favartia {Pygmaepterys) avatea Houart & Trôndlé, 2008 (holotype MNHN 20172); 8. Favartia {Pygmaepteiys) lifouensis (holotype MNHN 24174).

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NoVAREX 14(2): 43-48, 10 juin 2013

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R. II0UART& M. Shvhrns

New species of Favartia (Pygmaepterys) from Hawaii.

Etyinology. This species is named for Mr. Kern Osterstock, nicknamed "Kerno", in récognition of his générons support and help in developing a System allowing the junior author to dredge to the depths that this species has been shown to inhabit.

Remarks. From the sixteen Indo-West Pacific species, three hâve a last teleoconch whorl with only 3 axial varices and need not be compared further here: F. {P.) cracentis (Houart, 1996) from Indonesia, F. {P.) dondani (Kosuge, 1984) from the Philippines and F. {P.) menoiii (Houart, 1990) from New Caledonia. Six species are confined to the Indian Océan, namely F. (P.) alfredensis (Bartsch, 1915) from Port Alfred, South Africa, F. {P.) maraisi (Vokes, 1978) from Northern Transkei, South Africa, F. (P.) isabelae Houart & Rosado, 2008 from Madagascar and Mozambique, F. {P.) adenensis (Houart & Wranik, 1989), F. (P.) yemenensis (Houart & Wranik, 1989) and F. (P.) paulboschi Smythe & Houart, 1984 ail from the Gulf of Aden, Yemen and Oman. Ail these species are very different from F. (P.) kernoi and need not be compared here either.

The remaining seven species are F. {P.) civatea Houart & Trôndlé, 2008 from the Austral Archipelago and New Caledonia, F. {P.) bellini (D'Attilio & Myers, 1985) from the Philippines and Okinawa, F. (P.) circinata Houart & Héros, 2012, from Taiwan, F. (P.) funafiitiensis (Hedley, 1899) from the Funafuti Atoll, F. (P.) kurodai Nakamigawa & Habe, 1964 from Japan, F. {P.) lifouensis Houart & Héros, 2012 from New Caledonia and F. (P.) philcloveri (Houart, 1984) from the Philippines.

The Shell characters of Favartia (P.) circinata, F. {P.) kurodai and F. (P.) philcloveri differ markedly from those of F. (P.) kernoi n. sp.

Favartia (P.) kernoi n. sp. was erroneously identified as F. {P.) funafiitiensis, starting from Kaicher (1979) who illustrated a shell from Pokai Bay, Oahu, Hawaii. The same specimen was illustrated by Vokes &

D’Attilio (1980) and was then also identified as A [P] funafiitiensis, which following the authors only differs in having several small denticles on the inner lip. Another specimen of A {P.) kernoi n. sp., this one from the Philippines, from the Don Pisor collection was illustrated by D’Attilio & Myers (1985a and 1985b), also as F. (P) funafiitiensis. The locality of this last specimen seems very doubtful as no other specimen has been signalized from the Philippines since then, in spite of the extensive dredging that has been performed there since the end of the seventies. The label could hâve been mixed with that of another species.

The holotype of Favartia funafiitiensis (Figs 15-16) is a small specimen with 5 teleoconch whorls. It is probably immature, the protoconch is conical with 2+ protoconch whorls (Fig. 5), probably more than 3 (First whorls missing in the holotype) with an abapical keel on the last whorl and a sinusigeral notch, denoting a planktotrophic larval development. The last teleoconch whorl bears 6 varices and 5 low primary cords (P1-P5). PI is the strongest, P2 and P3 are slightly weaker, similar in size, P4 is smaller, P5 is the smallest. The shoulder is smooth except for axial lamellae. The siphonal canal is short and broadly open. The aperture is small, the columellar lip smooth except for 3 small knobs abapically. There are 4 denticles within the outer lip (D1-D4), DI is the strongest denticle.

Favartia funafiitiensis resembles F. philcloveri from the Philippines (Fig. 23). However, F. philcloveri is larger (with one supplementary teleoconch whorl), has 6 obvions primary spiral cords on the last whorl (Pl- P6), secondary cords, and numerous obvions lirae, including 7 or 8 on shoulder. P2, P3 and P4 are the strongest, almost similar in size, PI is weakly narrower. P5 and P6 are the smallest. The aperture of F. philcloveri is comparatively larger with a small pariétal tooth, a nan'ow, obvions ID and D1-D5 (D4- D5 fused).

Figures 9-23

9-14. Favartia (Pygmaepterys) kernoi n. sp.

9-13. Hawaii, dredged off Kihei, Maui, 20°39.180" N, 156°28.598" W to 20 38.650" N, 156 30.494" W, 170 m. 9-10. Holotype MNHN 25838, 10.7 mm; 11-12. Paratype RH, 10.5 mm; 13. Paratype IRSNB IG 32388/MT 2798 9.5 mm. 14. Hawaii, dredged off Kihei, Maui, 20°39.868" N, 156°29.347" W to 20°38.812" N, 156 28.733" W, 128 m, paratype MS, 13.3 mm.

15-16. Favartia (Pygmaepterys) funafiitiensis (Hedley, 1899), on W slope of Funafuti Atoll, Ellice Group, 8°29' S, 179°4' E, 73-146 m, holotype AMS C. 006004, 8.3 mm; 17-18. Favartia (Pygmaepteiys) bellini (DAttilio & Myers, 1985), Japan, Okinawa, Ryuku Is, 52 m, holotype SDNHM 83065, 9.9 mm; 19-20. Favartia (Pygmaepterys) avatea Houart & Trôndlé, 2008, French Polynesia, Austral archipelago, Rimatara, 22°38' S, 152°50' W, 250-280 m, holotype MNHN 20172, 1 1.2 mm; 21-22. Favartia (Pygmaepterys) lifouensis Houart & Héros, 2012, New Caledonia, Loyalty Islands, Lifou, Baie du Santal, Shclter Reef, 20°54' S, 167°02' E, 100-120, holotype MNHN 24174, 1 1.5 mm; 23. Favartia (Pygmaepteiys) philcloveri (Houart, 1984), Bohol, Philippines,

RH, 1 3.4 mm.

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Novapex 14(2): 43-48, lOjuin 2013

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New species of Favartia {Pygmaepterys) from Hawaii.

Both differ from F. kenwi n. sp. by their protoconch morphology and other shell characteristics.

Favartia (P.) avatea (Figs 7, 19-20) also resembles F. {P.) kenwi n. sp., however, the protoconch of F. avatea remains unknown. It differs in having a less shoLildered shell, a broader and more strongly sloping shoulder ramp, weakly lower spiral cords, a more obvions P6 cord and shorter spinelets at intersection of axial and spiral sculpture. F. (P.) avatea also differs in having more numerous axial varices: 9 on second whorl, 7 or 8 on third and fourth and 6 on last whorl. The aperture also differs in being larger and more narrowly ovate with a deeper, narrower anal notch and higher, stronger denticles within the outer lip, with D1-D2 fused (Figs 6 and 7).

Favartia {P.) lifouensis (Figs 8, 21-22) differs in having a higher spire, narrower spiral cords, with additional IP, si, s4, s5 and s6. The siphonal canal is comparatively broader and shorter. The apeiture also differs in being more strongly ovate with a narrower, deeper anal notch and in having more obvions inner apertural denticles (Figs 6 and 8).

Favartia {P.) bellini (Figs 17-18) has a more lightly built, narrower shell with a higher spire, narrower and more numerous spiral cords, occasionally with primary and secondary cords of same strength. The apertural denticles in F. {P.) bellini are lower and more numerous, occasionally split.

ACKNOWLEDGEiVlENTS

We are grateful to Virginie Héros (Muséum national d'Histoire naturelle, Paris, France) for the loan of the type material of Favartia {Pygmaepterys) avatea, to lan Loch and Janet Waterhouse (Australian Muséum, Sydney, Australia) for the loan of the holotype of Favartia {Pygmaeptetys) fwiafutiensis, to Margaret N. Dykens (San Diego Natural History Muséum of California, USA) for the loan of the holotype and paratype of Favartia {Pygmaepterys) bellini, to Pauline Fiene (Maui, Hawaii, USA) for her comments and corrections on the manuscript, to Thieiry Backeljau and the staff. Institut royal des Sciences

naturelles de Belgique for collaboration in many ways, and to Shirley Speer (Maui, Hawaii, USA) for pulling thousands of feet of rope and sorting sloshing dredge spoils on a rolling boat with unwavering dedication.

REFERENCES

DAttilio, A. & Myers, B.W. 1985a. A new species of Pygmaeptetys Vokes from the Western Pacific (Gastropoda: Muricidae). The Nautilus 99(1): 9-13. DAttilio, A. & Myers, B.W. 1985b. Illustrations of Recent Pygmaeptetys-, Muricopsinae. The Festiviis 27(6): 59-63.

Houart, R. & Rosado, J. 2008. Description of a new muricopsine species from Madagascar and Mozambique. Gloria Maris Al {\ -2)-. 1-7.

Houart, R. & Trôndle, J. 2008. Update of Muricidae (excluding Coralliophilinae) from French Polynesia with description of ten new species. Novapex 9(2- 3): 53-93.

Houart R. & Héros, V. 2012. New species of Muricidae (Gastropoda) and additional or noteworthy records from the western Pacific. Zoosystema 34( 1 ): 2 1 -37. Kaicher, S.D. 1979. Card catalogue of world-wide shells, Muricidae IV. Privately publ. St. Petersburg, Florida.

Merle, D. 2001 . The spiral cords and the internai denticles of the outer lip in the Muricidae: tenuinology and methodological comments. Novapex 2{3y. 69-91.

Merle, D. 2005. The spiral cords of the Muricidae (Gastropoda, Neogastropoda): importance of ontogenetic and topological correspondences for delineating structural homologies. Lethaia 38: 367- 379.

Vokes, E.H. & DAttilio, A. 1980. Pygmaeptetys, a newly described taxon of Muricidae (Mollusca: Gastropoda), with the description of three new species from the Cenozoic of the Western Atlantic. Tulane Studies in Geology and Paleontology.

16(2): 45-54.

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Les marées de 20 1 3

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Novapex / Société 14(2), 10 juin 2013

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NVVAPEX

/Swüil^

Prochaines activités de la SBM

Claude VILVENS

Lieu de réunion : A partir de 14h, à notre local habituel :

Salle "Memling” (1er étage - ascenseur) - Rue de Genève, 470b - Schaerbeek (Bruxelles)

SAMEDI 7 SEPTEMBRE 2013

Tout le monde : Atelier "Posters de la SBM"

On connaît les posters édités par la SBM : au nombre de 3, ils montrent respectivement les Terrestres de Wallonie, les Dulcicoles de Wallonie et les Clausiliidae du monde entier. Ces posters sont très appréciés des naturalistes et constituent pour la SBM une belle carte de visite. Aussi, nous envisageons d'en eréer de nouveaux, notamment "Coquillages du monde entier" (donc, un pour gastéropodes exotiques et un pour bivalves exotiques) et "Mollusques de Méditerranée" (mais ce ne sont que des exemples, toutes les idées seront les bienvenues dans la suite). Pour construire ees posters, nous allons travailler en "live" : nous vous demandons d'apporter certaines de vos meilleures photos de coquilles eorrespondant aux thèmes eités (bien sûr, des photos personnelles - pas des photos d'Internet protégées par copyright ;-) !). Nous construirons alors sur nos portables raccordés à un projecteur les posters souhaités. Un bon moyen de combiner connaissances et design ;-) ! Nous comptons sur vous !

SAMEDI 21 SEPTEMBRE 2013

Tout le monde : L*excursion d'automne de la SBM.

Cette fois, direction le Hainaut : nous allons prospeeter sur le site du terril du Six Perrier à Souvret ; *** Rendez-vous 9h30-10h à la gare de Courcelles ***

Comme d'habitude, les informations fraîches seront disponibles sur notre site Internet (http://www.societe- belge-de-malacologie.be/ Réunions). Comme d'habitude aussi, il convient de prévoir d'emporter sa bonne humeur, un guide de détermination (j'en connais des pas mal ;-)) ... et sans doute aussi bottes et vêtements de pluie (en principe, il fera eneore doux et ensoleillé, mais bon, on ne sait jamais ;-) ...).

SAMEDI 12 OCTOBRE 2013

Tout le monde : L'EXPOSITION ANNUELLE DE LA SBM.

Eh oui, voiei revenu le rendez-vous quasi-rituel de l'exposition de coquillages par les membres de la Société. Nous occuperons l'une des deux salles Memline ou Brueghel au même étage de notre adresse habituelle de réunion, selon les disponibilités (voir à nouveau sur notre site Internet).

Cette exposition est l'occasion pour chacun de montrer l'un ou l'autre aspeet de la malacologie qui lui tient à cœur. Aucune condition particulière n'est requise et tout le monde est cordialement invité à participer et aussi, bien sûr, à venir admirer quelques spécimens qui font la fierté de la collection de nos membres ! En particulier, pourquoi ne pas inviter quelques-uns de vos amis à venir découvrir notre passion ?

— PUIS : La tradition du banquet sera aussi respectée en tant qu'événement gastronomique attendu : nous vous proposons en effet de nous retrouver au traditionnel banquet annuel de la SBM oui débutera à 19h (voir annonce ci- dessous)

Réservez déjà dans vos agendas le 9/11/2012 et le 14/12/2012.

Pour les informations de dernière minute :

J Sur Internet :

■U http://www.societe-belge-de-malacologie.be/

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Novapex / Société 14(2), 10 juin 2013

Banquet de la Société Belge de Malacologie

le samedi 12 octobre 2013 à 19h

au restaurant ;

Le Phénix

Rue de Genève, 480 - 1030 Bruxelles

- donc juste à côté de la salle de réunion ! -

Nous vous emmenons à deux pas du lieu de réunion, au restaurant Le Phénix, lequel nous propose :

♦ Apéritif maison Ot/Prosecco

♦ Jambon de Parme et son melon Ot/Duo de fondants-croquette de fromage et crevettes

♦ Escalope du chef-jambon, fromage, gratinée OU Cabillaud à la dijonnaise

♦ Bavarois aux framboises

♦ Café

♦ 1/2 bouteille de vin par personne {rouge, blanc ou rosé}

Extra à payer individuellement en supplément.

Prix : 33,50 €

Il est impératif de réserver afin que le restaurateur puisse nous réserver le meilleur accueil

Comment réserver ?

Pour le 1er octobre 2013, au plus tard, il convient de virer la somme correspondant au nombre de menus réservés au compte BBL ; 310-1142433 — 53 de

Madame Annie Langleit, avenue Cicéron, 27/92 à 1 140 - Bruxelles (pas de paiement à la SBM, s’il vous plaît !)

Afin de préparer au mieux cette soirée, vous voudrez bien communiquer à Annie (annie.langleit@skynet.be), après la réservation, vos choix de plats (entrée et plat principal) - le restaurateur nous en sera très reconnaissant !

Nous nous réjouissons de vous rencontrer lors de cette joyeuse activité !

Bonnes vacances à tous !!!

Novapex/Société : la publication généraliste de la SBM

Rédacteurs en chef ; Claude Vilvens & Etienne Meuleman

Tous les articles généraux sont les bienvenus pour Novapex/Société © !

Afin de faciliter le travail de la Rédaction, il est vivement (et le mot est faible ;-)) souhaité de respecter les règles suivantes pour les articles proposés :

♦ document MS-Word (pour PC Windows XP ou 7);

♦ police de caractères Times New Roman;

♦ texte de taille 10, titres de taille 12;

♦ interligne simple;

♦ toutes les marges à 2,5 cm;

♦ document en une seule section;

♦ pas de mode colonne;

♦ photos en version électronique JPG ou PNG.

Merci pour les Scribes ;-) ! N'hésitez pas à demander une page avec en-tête pour cadrer au mieux vos travaux (vilvens.claude@skynet.be ou e.meuleman@skynet.be).

Novapex/ Société 14(2), 10 juin 2013

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Rapana venosa (Valenciennes, 1846) en Mer de Marmara

Christiane DELONGUEVILLE

Avenue Den Doom, 5 — B - 1180 Bruxelles - christiane.delongueville@skvnet.be

Roland SCAILLET

Avenue Franz Guillaume, 63 — B - 1140 Bruxelles - scaillet.roland@skvnet.be

MOTS-CLEFS Rapana venosa - Mollusques épibiontes - Mollusques parasites - Mer de Marmara KEY-WORDS Rapana venosa - Epibiontic molluscs - Parasitic molluscs - Marmara Sea

RÉSUMÉ

Sur des gros spécimens de Rapana venosa (Valenciennes, 1846) (Muricidae) - récoltés en Mer de Marmara - couverts d’algues, de concrétions calcaires et de vers, on peut identifier des mollusques épibiontes comme Mytilaster lineatus (Gmelin, 1791) (Mytilidae) présents en grand nombre. Acanthochitona fascicularis (Linnaeus, 1767) et Chiton olivaceus Spengler, 1797 (Polyplacophora) sont occasionnellement fixés dans l’ombilic de ces gastéropodes. Le test très épais et déjà endommagé par l’action perforante d’éponges est creusé de galeries et d’alvéoles abritant des mollusques bivalves perforants comme Petricola lithophaga (Philippson, 1788) (Petricolidae) et Gastrochaena dubia (Pennant, Mil) (Gastrochaenidae). Ces derniers peuvent être considérés comme parasites lorsque leur nombre met en danger l’intégrité de la coquille en la perforant de part en part.

ABSTRACT

On large specimens of Rapana venosa (Muricidae) - collected in the Marmara Sea - covered with algae, calcareous concrétions and worms, one can identify a large number of epibiontic molluscs like Mytilaster lineatus (Gmelin, 1791) (Mytilidae). Aeanthochitona fascicularis (Linnaeus, 1767) and Chiton olivaceus Spengler, 1797 (Polyplacophora) are occasionally fixed inside the umbilicus of the gastropods. The thick shell, already damaged by the action of perforating sponges, is carved into galleries and cells harbouring perforating bivalve molluscs like Petricola lithophaga (Philippson, 1788) (Petricolidae) and Gastrochaena dubia (Pennant, 1777) (Gastrochaenidae). These can be considered parasitic when their number endangers the integrity of the shell by perforating it ail the way through.

INTRODUCTION

La Mer de Marmara établit le lien entre la Méditerranée et la Mer noire. La salinité de ses eaux est intermédiaire entre celles des deux mers limitrophes (Méditerranée 38 %o - Mer Noire 17-18 %o). Le nombre d’espèces de mollusques qui y vit est moindre qu’en Méditerranée mais plus important qu’en Mer Noire (Oberling 1969-1971). Une population de Rapana venosa (Valenciennes, 1846), Muricidae invasif originaire de l’Océan Pacifique, y est implantée depuis de nombreuses années (Çinar et al. 2005). Rapana venosa atteint des tailles considérables (jusqu’à 18 cm) et les spécimens gérontiques sont souvent encroûtés et associés à d’autres mollusques. La présence d’un petit Mytilidae - Mytilaster lineatus (Gmelin, 1791) - avait été signalée dans l’ombilic d’un individu récolté en Mer Noire (Anonyme 1990).

RÉCOLTES PERSONNELLES

En mai 2012, un lot d’une vingtaine de spécimens vivants de Rapana venosa (Valenciennes, 1846) a été acquis auprès d’un pêcheur à Marmara Ereglisi (Figs. 1 et 3) en Mer de Marmara (Turquie). Les gastéropodes ont été capturés au filet sur un fond sablo-fangeux entre six et dix mètres de profondeur. Les plus grands individus mesuraient de 1 5 à 1 6 cm. Les coquilles étaient couvertes d’algues, de balanes, de bryozoaires encroûtants, de vers tubicoles {Spirobranchus sp.) et d’éponges perforantes. Cette multiplicité d’épibiontes et de parasites était loin de donner un aspect attrayant aux coquilles (Figs. 4 - 5). Des spécimens récoltés 22 ans plus tôt en eaux plus profondes (30 à 40 mètres) au large de Tekirdag (Delongueville & Scaillet 2007) étaient quant à eux en parfait état de conservation et dépourvus d’épibiontes (Fig. 2). Si les individus collectés à Marmara Ereglisi n’étaient pas esthétiquement attrayants, ils avaient le mérite d’être très intéressants, car ils ont permis d’observer la présence de nombreux autres mollusques qui avaient élu domicile sur la coquille parmi les concrétions et même dans l’épaisseur du test.

Mytilaster lineatus (Gmelin, 1791) (Fig. 9) est un petit Mytilidae (15 à 20 mm) commun en Méditerranée, le long des côtes italiennes, grecques et turques, et en Mer Noire. Les spécimens étaient fermement fixés dans l’ombilic des Rapana, mais aussi entre les plis de la coquille, parmi les tubes calcaires des Spirobranchus ou même à l’intérieur de balanes vides (Figs. 6 et 7). Sur un spécimen haut de 14 cm (Fig. 6), il y